Olegia dorotheae ( Hoernes & Auinger, 1891 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 83-85

Olegia dorotheae ( Hoernes & Auinger, 1891) nov. comb.

Figs 25C 1 –C 3 , D 1 –D 3 , E 1 –E 3 , F 1 –F 3, 3 Ae, 5, 7

Clavatula Dorotheae —Hoernes 1891: 128 [nomen nudum].

* Pleurotoma (Clavatula) Dorotheae nov. form. — Hoernes & Auinger 1891: 336, pl. 46, figs 5–7 [non fig. 4, = ‘ Clavatula ’ mariae ( Hoernes & Auinger, 1891) ].

Pleurotoma conf. baccifera Bell. — Vetters 1910: 148 [non Olegia baccifera ( Bellardi, 1877) ].

Clavatula dorotheae (Hörnes & Auinger, 1891) —Harzhauser 2002: 118, pl. 10, figs 10–11, [non figs 12–13, = ‘ Clavatula ’ mariae ( Hoernes & Auinger, 1891) ].

non Clavatula (Clavatula) dorotheae R. Hoernes et Auinger — Švagrovský 1958: 14, pl. 4, fig. 1 [= Olegia sp. ].

non Clavatula (Clavatula) dorotheae R. Hoernes et Auinger — Švagrovský 1960a: 102, pl. 13, fig. 1 [= Olegia sp. ].

non Pleurotoma (Clavatula) dorotheae R. Hoernes et Auinger — Florei 1961: 679, pl. 6, fig. 34 [= belongs to Perrona perron group].

Type material. Lectotype designated herein: NHMW 1949 /0005/0040a, SL: 18.2 mm, MD: 7.4 mm, Kleinebersdorf ( Austria), illustrated in Hoernes & Auinger (1891, pl. 46, figs 5a–c), figs 25C 1 –C 3 . Paralectotypes: NHMW 1949 /0005/0040b, SL: 12.9 mm , MD: 5.4 mm, Kleinebersdorf ( Austria), illustrated in Hoernes & Auinger (1891, pl. 46, figs 7a–c), figs 25D 1 –D 3, 3 Ae ; NHMW 1949 /0005/0040c, SL: 17.7 mm , MD: 7.3 mm, Kleinebersdorf ( Austria), illustrated in Hoernes & Auinger (1891, pl. 46, figs 6a–c), figs 25E 1 –E 3 .

NHMW 2021/0146/0001, SL: 15.7 mm, MD: 6.6 mm, Kleinebersdorf ( Austria), figs 26F 1 –F 3.

Additional studied material. 25 spec. NHMW 1976 /1784/0002, Kleinebersdorf ( Austria) , 2 spec., NHMW 1949 /0005/0040, Karnabrunn ( Austria); NHMW 1861 /0050/00148 , 12 spec., Kleinebersdorf ( Austria); NHMW 1859 /0045/0094 , 2 spec., Niederkreuzstetten ( Austria) .

Revised description. Shell small, solid, moderately broad fusiform; apical angle ~40–47°. Protoconch not preserved. Teleoconch of at least nine whorls (earliest whorls not preserved). Early teleoconch whorls with broad, smooth subsutural cord and beaded suprasutural cord, separated by concave mid-portion. Sculpture changing on fourth teleoconch whorl; subsutural cord swelling into prominent, rounded subsutural collar with poorly defined tubercles marking whorl periphery, mid-portion concave, narrow, suprasutural row of beads well developed. Entire surface covered in fine spiral threads, overrunning collar and beads. Suture narrowly impressed, weakly undulating. Last whorl 65% of total height, barrel shaped; subsutural collar prominent, rounded, subsutural ramp narrow, concave. Shoulder weakly developed, delimited by row of large poorly defined, axially elongated tubercles that extend axially to form vague ribs; convex below, moderately constricted at base. Siphonal fasciole poorly delimited, twisted. Below shoulder primary cords not clearly defined, covered in irregular, flattened, poorly defined cords extending to fasciole. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately deep and wide, asymmetrically U-shaped, with apex mid-ramp. Siphonal canal moderately short and narrow, straight to slightly deflected to the left, shallowly notched at tip. Columella straight to weakly convex, smooth. Columellar and parietal callus thickened, forming broad callus rim.

Discussion. Olegia dorotheae ( Hoernes & Auinger, 1891) was partly confused with ‘ Clavatula ’ mariae by Hoernes & Auinger (1891) and Harzhauser (2002), but ‘C. ’ mariae has a conical spire, a weaker subsutural collar and lacks axial ribs on the last whorl. Olegia dorotheae is reminiscent of O. brigittae ( Hoernes & Auinger, 1891) , but differs in its swollen subsutural collar and more concave whorls. ‘ Clavatula ’ geniculata Bellardi, 1877, from the Langhian of Albugnano ( Italy), develops similar sculpture, but differs in its high spire and more strongly constricted base (see Bellardi 1877: 174, pl. 5, fig. 37). The specimen from the middle Miocene of Albania, described by Martelli & Nelli (1910: 550, pl. 11, fig. 5) as ‘ C. ’ geniculata is unrelated with that species and differs from ‘ C. ’ geniculata and O. dorotheae by the conical spire and the absence of any nodes or beads. The single specimen described by Švagrovský (1959, 1960a) from the Badenian of Kuzmice ( Slovakia) as C. dorotheae differs from the Karpatian type in its constricted and more angulated base and longer siphonal canal and lacks the prominent subsutural collar.

Paleoenvironment. Estuarine, littoral mud flats ( Zuschin et al. 2004). Distribution in Central Paratethys. Karpatian (early Miocene): Korneuburg Basin: Kleinebersdorf, Karnabrunn, Obergänserndorf ( Austria) (Harzhauser 2002, 2003); North Alpine-Carpathian Foreland Basin: Niederkreuzstetten ( Austria) (Harzhauser 2002, 2003).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 25. A1–A3. Olegia brigittae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0144/0001, Lysice (the Czech Republic). B1–B3. Olegia brigittae (Hoernes & Auinger, 1891), paralectotype, NHMW 2021/0145/0001, Niederleis (Austria). C1–C3. Olegia dorotheae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0040a, Kleinebersdorf (Austria). D1–D3. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). E1–E3. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW NHMW 1949/0005/0040c, Kleinebersdorf (Austria). F1–F3. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 2021/0146/0001, Kleinebersdorf (Austria).