Megaclavatula subsculpta (Schaffer, 1912) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 75-76

Megaclavatula subsculpta (Schaffer, 1912) nov. comb.

Figs 22B 1 –B 3

* Pleurotoma (Clavatula) asperulata Lam. var. subsculpta Schff. — Schaffer 1912a: 134, pl. 49, figs 9–10.

Pleurotoma (Clavatula) asperulata Lam. var. subsculpta Schff. — Schaffer 1912b: 327.

C [lavatula]. (C [lavatula]) asperulata subsculpta Schff. — Sieber 1958a: 157.

Clavatula (Clavatula) asperulata subsculpta Schaffer — Mandic & Steininger 2003: 274.

Type material. Lectotype designated herein: KM Inv. No. F /1125, Maria Dreieichen ( Austria), SL: 28.5 mm, MD: 10.6 mm, illustrated in Schaffer (1912a, pl. 49, fig. 9), figs 22B 1 –B 3 . Paralectotype: KM Inv. No. F /1126, Maria Dreieichen ( Austria), illustrated in Schaffer (1912a, pl. 49, fig. 10). The specimen is lost .

Revised description. Shell medium-sized, solid, moderately broad fusiform with high conical spire; apical angle 34°. Protoconch and early teleoconch whorls not preserved. Spire whorls with moderately swollen, indistinctly beaded subsutural collar, weakly concave mid-whorl, and prominent, coarsely beaded suprasutural cord. Suture narrowly but deeply incised, undulating below beads. Last whorl 55% of total height, poorly delimited subsutural collar with ten weakly pointed tubercles, concave subsutural ramp, roundly angled at shoulder and peribasal cord. Base moderately constricted, concave. Siphonal fasciole poorly delimited, of moderate length, slightly twisted. Three primary spiral cords, adapical with large rounded, slightly opisthocline tubercles delimiting shoulder, abapical narrower delimiting base, third intercalated at mid-whorl of similar width to peribasal cord. Weaker irregular cords over base and siphonal fasciole. Close-spaced axial growth lines cover surface.Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus deep, moderately wide, asymmetrically V-shaped, with apex below subsutural collar in deepest part of subsutural concavity. Siphonal canal moderately long, narrow, slightly deflected to the left, weakly notched at tip. Columella weakly excavated, slightly twisted at fasciole, smooth. Columellar and parietal callus thin, adherent, poorly delimited from the base.

Discussion. Schaffer (1912a) established this species as subspecies of Megaclavatula asperulata ( Lamarck, 1822) , which differs by its more delicate sculpture on the early teleoconch whorls, the presence of prominent spiral cords on the spire whorls, spinose tubercles on the subsutural collar, and a shorter last whorl. In view of the poor preservation of the early whorls, placement in Megaclavatula is tentative and will need confirmation.

Paleoenvironment. The occurrence at Maria Dreieichen ( Austria) suggests shallow inner neritic to lagoonal environments (own data).

Distribution in Central Paratethys. Eggenburgian (early Miocene): North Alpine-Carpathian Foreland Basin: Maria Dreieichen ( Austria) ( Mandic & Steininger 2003).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 22 A1–A3. Megaclavatula laevigata (Eichwald, 1830), NHMW 1846/0037/0308, Baden (Austria). B1–B3. Megaclavatula subsculpta (Schaffer, 1912), lectotype, KM Inv. No. F/1125, Maria Dreieichen (Austria). C1–C2. Megaclavatula nemethi (Kovács & Vicián, 2021), holotype, HNHM, PAL 2021.21.1, Letkés (Hungary). D. Megaclavatula nemethi (Kovács & Vicián, 2021), paratype, NHMW 2021/0002/0003, Letkés (Hungary). E1–E2. Megaclavatula? szontaghyi (Strausz, 1926), holotype, M.116, Bán, Baranya County (Hungary). F. Megaclavatula neudorfensis (Schaffer, 1898), holotype, NHMW 2021/0137/0001, Devínska Nová Ves (Slovakia). G1–G3. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). H1–H3. Megaclavatula pilleri nov. sp., paratype, NHMW 1949/0005/0051b, Baden-Sooss (Austria).