Megaclavatula evae ( Hoernes & Auinger, 1891 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 64-65

Megaclavatula evae ( Hoernes & Auinger, 1891) nov. comb.

Figs 21A 1 –A 3 , B 1 –B 3 , 5 , 6

Clavatula Evae —Hoernes 1891: 129 [nomen nudum].

* Pleurotoma (Clavatula) Evae nov. form.— Hoernes & Auinger 1891: 344, pl. 44, figs 3–4.

Pleurotoma (Clavatula) asperulata Lamk. var.– Hoernes & Auinger 1891: 343, pl. 44, figs 5a–b [non ‘ Clavatula ’ asperulata Lamarck, 1822 ].

Pleurotoma Clavatula asperulata Lam. — Sieber 1947: 157 (partim) [non Megaclavatula asperulata ( Lamarck, 1822) ].

Clavatula asperulata (Lk.) — Sieber 1953: 188 (partim) [non Megaclavatula asperulata ( Lamarck, 1822) ].

C [lavatula]. (C [lavatula]) evae (R. Hörn. et Au.) — Sieber 1958a: 157.

Clavatula asperulata var. evae Hoernes et Auinger , sp. 1891— Glibert 1960: 37.

non Clavatula cf. evae (R. Hoernes & Auinger, 1891) — Mikuž 1998: 74, pl. 4, fig. 2 [= abraded ‘ Clavatula ’ camillae ( Hoernes & Auinger, 1891)]. non Clavatula evae ( Hoernes & Auinger, 1891) — Bałuk 2003: 34, pl. 5, figs 1–8 [= Megaclavatula amaliae ( Hoernes & Auinger, 1891)]. non Clavatula cf. evae (R. Hoernes & Auinger, 1891) — Mikuž 2009: 32, pl. 10, fig. 141 [= abraded ‘ Clavatula ’ camillae ( Hoernes & Auinger, 1891)].

Type material. Lectotype designated herein: NHMW 1949 /0005/0047a, Guntersdorf ( Austria), SL: 69.9 mm, MD: 23.4 mm, illustrated in Hoernes & Auinger (1891, pl. 44, fig. 3), figs 21B 1 –B 3 . Paralectotype: NHMW 1949 /0005/0047b, Guntersdorf ( Austria), SL: 66.4 mm , MD: 23.3 mm, illustrated in Hoernes & Auinger (1891, pl. 44, fig. 4), figs 21A 1 –A 2.

Additional studied material. NHMW 1851/0026/0044 a, SL: 75.1 mm, MD: 25.3 mm, Grund ( Austria).

Revised description. Shell large, solid, slender fusiform with tall conical spire; apical angle~30–35°. Protoconch not preserved. Teleoconch of at least ten weakly concave whorls. Early teleoconch whorls with tripartite spiral sculpture; weak, vaguely beaded subsutural cord, slightly weaker, beaded mid-whorl cord, prominent strongly beaded suprasutural cord. Sculpture changing around fifth to sixth teleoconch whorl; whorls increase in height, subsutural cord develops into weak collar with wide spaced, low, spiny tubercles placed short distance from suture; mid-cord disappears. Beads on subsutural cord becoming low, wide spaced tubercles partly covered by subsequent whorl. Occasional indistinct spiral cord present at mid-whorl. Suture narrowly but deeply incised, weakly undulating. Last whorl ~57% of total height, poorly delimited. Weakly spinous subsutural collar. Very weakly concave subsutural ramp, rounded to slightly angular at weakly inflated shoulder. Hardly to weakly delimited shoulder. Base moderately constricted; spiral sculpture weak, with shoulder and peribasal cords slightly stronger and weakly nodular, further primary spiral cord at mid-whorl between shoulder and peribasal cords. Weaker cords over base and siphonal fasciole. Siphonal fasciole of medium length, moderately delimited, slightly twisted. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus wide, moderately deep, asymmetrically broad Vshaped, with apex on adapical portion of ramp; siphonal canal moderately long, slightly recurved, shallowly notched at tip. Columella shallowly excavated, weakly twisted, smooth. Columellar and parietal callus weakly thickened, forming broad callus rim.

Discussion. Bałuk (2003) confused Megaclavatula evae with M. polonica . However, the shape and sculpture of M. evae distinguish it immediately from the broader and strongly coronate M. polonica (Pusch, 1837) . The specimens that Bałuk (2003) described as M. evae , in contrast, are not conspecific with this species, and differ in their prominent spiral sculpture, which is not developed by M. evae . Therefore, the specimens described by Bałuk (2003, pl. 5, 4–7) should be transferred to Megaclavatula amaliae ( Hoernes & Auinger, 1891) . ‘ Clavatula ’ gracilis Erünal-Erentöz, 1958 , from the Serravallian of Turkey, seems to be a related species and has a comparable slender outline. It differs from M. evae mainly in its longer siphonal canal and the more prominent tubercles on the subsutural collar (see Erünal-Erentöz 1958: 95, pl. 14, figs 20–23, pl. 15, fig. 12; Landau et al. 2013: 284, pl. 47, figs 4a–c).

‘ Clavatula ’ turriculata ( Grateloup, 1846) from the Burdigalian of Saint-Paul ( France) is a morphologically similar species in the northeastern Atlantic fauna but has close-set beads at the abapical suture of late teleoconch whorls and very prominent spiral sculpture (see Peyrot 1931, pl. 5, figs 1–2). ‘ Clavatula ’ sequini ( Mayer, 1858) from the Burdigalian of Italy, is slightly similar in shape and reduced sculpture, but differs in the tiny beads at the abapical suture and the spiral sculpture (see Mayer 1858, pl. 11, fig. 10; Bellardi 1877: pl. 7. fig. 17).

Paleoenvironment. At the locality Grund fossiliferous channel fills, which formed in middle to outer neritic environments bear allochthonous assemblages uniting coastal-mudflat faunas with inner neritic ones ( Zuschin et al. 2005; Roetzel 2009). Preservation and color of the shells, suggest, that Megaclavatula evae belonged to the allochthonous shallow water fauna and not to the autochthonous deep-water fauna.

Distribution in Central Paratethys. Badenian (middle Miocene): North Alpine-Carpathian Foreland Basin: Guntersdorf, Grund ( Austria) ( Hoernes & Auinger 1891).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia).

FIGURE 21 A1–A3. Megaclavatula evae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0047b, Guntersdorf (Austria). B1–B3. Megaclavatula evae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0047a, Guntersdorf (Austria). C1–C3. Megaclavatula grunerti nov. sp., holotype, NHMW A1382, Enzesfeld (Austria), D1–D3. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). E1–E3. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035b, Korytnica (Poland). F1–F3. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland).