Tomellana dulaii, Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 146-147

publication ID

https://doi.org/ 10.11646/zootaxa.5123.1.1

publication LSID

lsid:zoobank.org:pub:036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A

persistent identifier

https://treatment.plazi.org/id/039487D1-FF13-FF3A-FFBA-FEA96C8BFE81

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Plazi (2022-03-30 20:52:14, last updated by Valdenar 2024-02-29 15:27:11)

scientific name

Tomellana dulaii
status

sp. nov.

Tomellana dulaii View in CoL nov. sp.

Figs 42C View FIGURE 42 1 –C View FIGURE 1 3 View FIGURE 3 , 4Z View FIGURE 4 , 5 View FIGURE 5 , 7 View FIGURE 7

Pleurotoma Borsonii Bast. View in CoL — Hauer 1837: 419 [non ‘ Clavatula View in CoL ’ borsonii de Basterot, 1825].

Pleurotoma (Clavatula) Borsoni Bast. — Hoernes & Auinger 1891: 359, pl. 47, figs 12a–c [non ‘ Clavatula View in CoL ’ borsonii (de Basterot, 1825)].

? Clavatula (Perrona) cf. borsoni Bast. — Csepreghy-Meznerics 1969: 97, pl. 6, fig. 3 [non ‘ Clavatula View in CoL ’ borsonii (de Basterot, 1825)].

? Clavatula (Perrona) cf. borsoni Bast. — Csepreghy-Meznerics 1972: 32, pl. 15, fig. 26 [non ‘ Clavatula View in CoL ’ borsonii (de Basterot, 1825)].

non Clavatula (Perrona) semimarginata borsoni ( Basterot, 1825) — Hölzl 1958: 265, pl. 21, fig. 18 [= not a Clavatulidae View in CoL ].

non Clavatula (Perrona) semimarginata cfr. borsoni (Basterot) — Švagrovský 1958: 20, pl. 6, fig. 6 [= Perrona lydiae ( Hoernes & Auinger, 1891) View in CoL ].

Type material. Holotype: NHMW 2021 View Materials /0148/0002, SL: 45.3 mm, MD: 15.9 mm, Nemeşeşti ( Romania), illustrated in Hoernes & Auinger (1891, pl. 47, figs 12a–c), figs 42C 1 –C 3, 4Z.

Type locality. Nemeşeşti ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, early/middle Badenian (Langhian).

Etymology. In honor of Alfréd Dulai (Hungarian Natural History Museum, Budapest), in recognition of his contributions to the paleontology of the Paratethys Sea.

Diagnosis. Moderately large, moderately broad fusiform with high last whorl and conical spire of weakly concave later flat-sided whorls. Narrow, carinate subsutural collar on last whorl. Almost flat subsutural ramp. Inflated mid-whorl with shoulder and ramp poorly delimited. Long siphonal canal. Spiral sculpture prominent only over base and fasciole.

Description. Shell moderately large, moderately broad fusiform with high last whorl and conical spire; apical angle 35°. Protoconch not preserved. Teleoconch of at least eight whorls. Early teleoconch whorls with narrow subsutural cord, beaded suprasutural cord, broad concave mid-portion bearing fine cords. On third whorl suprasutural cord of beads disappears, entirely obscured by succeeding whorl. Abapically subsutural collar remains narrow, weakly carinate, poorly delimited. Mid-whorl portion becomes less concave, straight-sided on penultimate whorl. Suture narrowly impressed, superficial, linear. Entire surface covered by fine cords. Suture narrowly impressed, linear. Last whorl 70% of total height. Subsutural collar narrow, carinate. Subsutural ramp broad, almost flat-sided. Shoulder poorly-delimited, rounded; mid-whorl inflated. Base weakly angled, strongly constricted. Siphonal fasciole poorly developed, weakly rounded. Sculpture of spiral threads over collar and ramp, mid-whorl smooth, narrow cords of alternating strength appearing on base and fasciole. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately wide and deep, asymmetrically U-shaped, with apex mid-ramp. Siphonal canal long, moderately wide, straight, shallowly notched at tip. Columella straight, smooth. Columellar and parietal callus thin, adherent.

Discussion. This species was identified by Hoernes & Auinger (1891) as Perrona borsoni (de Basterot, 1825) , originally described from the early Miocene of France (de Basterot 1825: 64, pl. 3 fig. 2). Perrona borsoni is currently treated as subjective junior synonym of Tomellana semimarginata ( Lamarck, 1822) ( Lozouet et al. 2001) , and differs from the Paratethyan species in its opisthocline axial riblets on the early teleoconch whorls ( Lozouet 2017, pl. 33, fig. 6), the higher spire and the rounded and less prominent subsutural collar. Perrona burdigalensis ( Peyrot, 1931) , from the early Miocene of France, is slightly reminiscent of Tomellana dulaii nov. sp., but its subsutural collar is even more strongly carinate, and it has a much higher spire (see Peyrot 1931, pl. 6, figs 30–31) (early teleoconch whorls of a specimen in the RGM collection place this species in Perrona ). Tomellana dulaii is close to Tomellana jouannetii ( Des Moulins, 1842) in general shape and sculpture, but the French early Miocene species has a convex subsutural collar and a higher spire (see Peyrot 1931, pl. 8, figs 43, 61, 62, 66, 71).

A specimen from the Badenian of Borsodbóta ( Hungary) described by Csepreghy-Meznerics (1969; 1972) might represent this species as well, but has an even wider apical angle and a shorter siphonal canal, which might be incomplete.

Paleoenvironment. Inner neritic environments based on the assemblage from Nemeșești ( Romania) (own data).

Distribution in Central Paratethys. Badenian (middle Miocene):? Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1969, 1972); Făget Basin: Nemeșești ( Romania) ( Hoernes & Auinger 1891).

Basterot, B. de (1825) Memoire Geologique sur les Environs de Bordeaux. Premiere partie comprenant les observations generales sur les mollusques fossiles, et la description particuliere de ceux qu'on rencontre dans ce bassin. Memoires de la Societe d'Histoire Naturelle de Paris, 2, 1 - 100. [https: // www. biodiversitylibrary. org / item / 23524 # page / 12 / mode / 1 up]

Csepreghy-Meznerics, I. (1969) Nouvelles gasteropodes et lamellibranches pour la faune hongroise des gisements tortoniensinferieurs de la Montagne de Bukk. Annales Historico-Naturales Musei Nationalis Hungarici Pars Mineralogica et Palaeontologica, 61, 63 - 127.

Csepreghy-Meznerics, I. (1972) La faune tortonienne-inferieure des gisements tufiques de la Montage de Bukk: Gastropodes II. Egri Muzeum Evkonyvebol (Annales Musei Agriensis), 8, 26 - 46. https: // library. hungaricana. hu / en / view / MEGY _ HEVE _ agria 08 _ 09 /? pg = 27 & layout = s

Des Moulins, M. Ch. (1842) Revision de quelques especes de Pleurotomes. Actes de la Societe linneenne de Bordeaux, 12, 109 - 185. [https: // www. biodiversitylibrary. org / item / 101660 # page / 164 / mode / 1 up]

Hauer, J. (1837) Uber das Vorkommen fossiler Thierreste im tertiaren Becken von Wien, und angehangte Vergleichung derselben mit den Uberresten derer gleichzeitigen Ablagerungen, nebst einigen nachtraglichen Bemerkungen uber das Maynzer- Becken von H. G. Bronn. Neues Jahrbuch fur Mineralogie, Geognosie, Geologie und Petrefaktenkunde, 1837, 408 - 431. [https: // www. biodiversitylibrary. org / item / 44261 # page / 428 / mode / 1 up]

Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]

Holzl, O. (1958) Die Mollusken-Fauna des oberbayrischen Burdigals. Geologica Bavarica, 38, 1 - 348.

Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]

Lozouet, P., Lesport, J. - F. & Renard, P. (2001) Revision des Gastropoda (Mollusca) du Stratotype de l'Aquitanien (Miocene inf.): site de Saucats ' Lariey', Gironde, France. Cossmannia, Hors-Serie 3, 1 - 190.

Lozouet, P. (2017) Les Conoidea de l'Oligocene superieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana, 19, 3 - 179.

Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]

Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56.

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FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

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FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

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FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

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FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

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FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

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FIGURE 42 A1–A3. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). B1–B2. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). C1–C2. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). D1–D2. Tomellana praecursor (Schaffer, 1912), lectotype, KM Inv. No. F/1129, Maigen (Austria). E. Tomellana praecursor (Schaffer, 1912), paralectotype, KM Inv. No. F/1130, Maigen (Austria). F. ‘Clavatula’? czarnockii (Bałuk, 2003), holotype. Catalogue No. 28.II. 179, Korytnica (Poland). F1–F3. Tomellana aueri nov. sp., holotype, NHMW 1874/0033/0009a, Bad Vöslau (Austria). G1–G3. Tomellana aueri nov. sp., paratype, NHMW 1874/0033/0009b, Bad Vöslau (Austria). H. Tomellana aueri nov. sp., paratype, NHMW 2013/0078/0436, Baden (Austria).

MD

Museum Donaueschingen

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Order

Neogastropoda

Family

Clavatulidae

Genus

Tomellana