Stenosagola pseudogracilis Park and Carlton

Park, J. - S. & Carlton, C. E., 2013, A Revision of the New Zealand GenusStenosagolaBroun, 1921 (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin 67 (3), pp. 335-359 : 341-343

Stenosagola pseudogracilis Park and Carlton , new species

Type Material. Holotype. NEW ZEALAND: WELLINGTON ( WN): ♂ (winged male), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: WN: Tararua Forest Park, Judd Ridge, Field’ s Track, Field Hut vic., 855m, 40″54.474′S 175″15.371′E, 24XI-26 XI 2005, broadleaf (Notho. meziesii-Weinmannia racemosa)- podocarp forest; FMHD#2005-040, berl., moss on log, A. Solodovnikov and D. Clarke; ANMT site 1153”, “ HOLOTYPE Stenosagola pseudogracilis Park and Carlton des. 2013”. Paratypes (n = 37: 30 males, 7 females). NEW ZEALAND: BAY OF PLENTY ( BP): 1♂, Kaimai-Mamaku Forest, Mt. Te Aroha, upper end Tui Mine Track, nr. Summit road, 775m, 37°31.685′S 175°44.684′E, 19 XI–26 XII 2005, low Nothofagus menziesii forest w/ Astelia ground layer, FMHD #2005-019, flight intercept trap, A. Newton and M. Thayer, ANMT site 1145 ( FMNH); 1♂, Kaimai-Mamaku Forest, Mt. Te Aroha, upper end Tui Mine Track, nr. Summit road, 775m, 37°31.685′S 175°44.684′E, 19 XI 2005, low Nothofagus menziesii forest w/ Astelia ground layer, FMHD #2005-021, berl., leaf and log litter, A. Newton and M. Thayer, ANMT site 1145 ( FMNH); GISBORNE ( GB): 2♂♂, Urewera N. P., Maungapohatu Rd., 3.2km E Taupeupe Saddle, 1110m, 38°36.975′S 177°02.753′E, 22 XI 2005, mossy Nothofagus menziesii forest, FMHD #2005-027, berl., leaf and log litter, A. Newton and M. Thayer, ANMT site 1148 ( FMNH); 2♂♂ (1♂,slide-mounted), Urewera N. P., Maungapohatu Rd., 3.2km E Taupeupe Saddle, 1110m, 38°36.975′S 177°02.753′E, 22 XI–23 XII 2005, mossy Nothofagus menziesii forest, FMHD #2005-025, flight intercept trap, M. Thayer and A. Solodovnikov, ANMT site 1148 ( FMNH); HAWKES BAY ( HB): 1♂, Kaweka Ra., Makahu Hut, 976m, 24 II 1971, J.I. Townsend, A.C. Eyles, litter 71/21 ( NZAC); MARLBOROUGH ( MB): 1♂ 1♀, Titirangi, 20 X 1969, F.A. Alack, litter 69/174 ( NZAC); 1♂, Mt. Robinson, S. Ridge, 518m, Kenepuru Hd., 13 III 1970, J.I. Townsend, litter 70/142 ( NZAC); 1♂, Okiwi Bay, 27 VIII 1965, J.I. Townsend, moss 65/440 ( NZAC); 1♂ 2♀♀, Head Fabians Valley, 23 X 1963, J.I. Townsend, litter 63/38 ( NZAC); MARLBOROUGH SOUNDS ( SD): 9♂♂ (1♂, slide-mounted), Opouri Saddle, above Tennyson Inlet, 540m, 15 XII 1984 – 5 I 1985, Nothofagus forest, A. Newton and M. Thayer 708, window trap ( FMNH); 5♂♂ 3♀♀ (1♂ 1♀, slide-mounted), Opouri Saddle, above Tennyson Inlet, 540m, 15 XII 1984 – 5 I 1985, Nothofagus forest, A. Newton and M. Thayer 708, berl., leaf and log litter, forest floor ( FMNH); 1♂, Opouri Saddle, above Tennyson Inlet, 540m, 15 XII 1984 – 5 I 1985, Nothofagus spp. forest A. Newton and M. Thayer 708 ( FMNH); 1♂, Tennyson Inlet, east side Duncan Bay, 30m, 15 XI 1984 – 5 I 1985, podo.- Nothofagus forest A. Newton and M. Thayer 709 ( DSC); 1♂, 70 km NE Nelson, Tennyson Inlet, 480m, 27 V 1982, FMHD #82-604, Beech forest litter, S. Peck ( FMNH); WELLINGTON ( WN): 3♂♂, Tararua Forest Park, Judd Ridge, Field’ s Track, Field Hut vic., 855m, 40° 54.474′S 175°15.371′E, 24 XI 2005, broadleaf (Notho. meziesii-Weinmannia racemasa)-podocarp forest, FMHD #2005-037, berl., sedge-base litter, A. Solodovnikov and D. Clarke, ANMT site 1153 ( FMNH).

Etymology. The specific epithet refers to the superficial similarity of this species to S. gracilis .

Diagnosis. Specimens of S. pseudogracilis are similar in appearance to those of S. gracilis , S. clarkei , S. ramsayi , S. domettensis , and S. eylesi but can be distinguished by the shape of the male genitalia ( Fig. 4a ). Members of this species can be separated from those of S. egmontensis and S. stewartensis by the glossy dorsal surface of the head ( Fig. 1d–f ).

Description. Length 1.6–1.8 mm ( Fig. 1d–f ). Head: Winged male head approximately 1.37 times, wingless male 1.37 times, female 1.39 times longer than wide. Winged male eye approximately 0.81 times, wingless male 0.67 times, female 0.35 times as long as temple. Winged male antennomere 3 subquadrate, 4 longer than wide, 5 subquadrate. Wingless male antennomere 3 slightly transverse, 4–5 subquadrate. Female antennomere 3 transverse, 4 subquadrate, 5 transverse. Thorax: Pronotum approximately 1.10 times longer than wide. Winged male elytra approximately 1.66 times, wingless male 1.35, female 1.14 longer than pronotum. Winged male meso- and metaventrites trapezoidal, approximately 1.48 times, wingless male 1.20, female 1.12 longer than pronotum. Aedeagus: Right apical lobe of male genitalia longer than left ( Fig. 4a ). Left paramere broader than right with setae present at tip and middle ( Fig. 4a ).

Distribution. BP (Bay of Plenty), HB (Hawkes Bay), MB (Marlborough), SD (Marlborough Sounds), WN (Wellington), WO (Waikato), New Zealand ( Fig. 3 , circles)

Habitat. Specimens of this species were collected with window traps, flight intercept traps, and sifting moss or leaf litter in Nothofagus , broadleaf, and podocarp forests.

Figures

Fig. 4. Stenosagola species. a) S. pseudogracilis, aedeagus, dorsal view, b) S. clarkei, aedeagus, dorsal view, c) S. egmontensis, aedeagus, dorsal view, d) S. ramsayi, e) S. egmontensis, winged male head, dorsal view; f) S. egmontensis, prothorax, ventral view; g) S. egmontensis, prothoracic leg, h) S. domettensis, aedeagus, dorsal view, i) S. eylesi, aedeagus, dorsal view, j) S. stewartensis, aedeagus, dorsal view. Scale bars = 0.1 mm.

Fig. 1. Habitus of Stenosagola species, dorsal view. a) Winged male of S. gracilis, b) Wingless male of S. gracilis, c) Female of S. gracilis, d) Winged male of S. pseudogracilis, e) Wingless male of S. pseudogracilis, f) Female of S. pseudogracilis, g) Wingless male of S. clarkei, h) Wingless male of S. egmontensis, i) Female of S. egmontensis, j) Winged male of S. ramsayi, k) Wingless male of S. ramsayi, l) Female of S. ramsayi, m) Wingless male of S. domettensis, n) Female of S. domettensis, o) Wingless male of S. eylesi, p) Wingless male of S. stewartensis. Scale bars = 1 mm.

Fig. 3. Locations where Stenosagola gracilis (triangles) and Stenosagola pseudogracilis (circles) have been collected in New Zealand.