Cyclocephala santandereana Neita, 2021

Neita-Moreno, Jhon C., 2021, A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia, Zootaxa 5026 (1), pp. 1-58 : 12-13

publication ID	https://doi.org/ 10.11646/zootaxa.5026.1.1
publication LSID	lsid:zoobank.org:pub:07E0C922-6B0F-4916-85C2-AD95146A8F1E
DOI	https://doi.org/10.5281/zenodo.10240010
persistent identifier	https://treatment.plazi.org/id/03932D70-B628-C37B-FF7B-08C5A1D0DEBB
treatment provided by	Plazi
scientific name	Cyclocephala santandereana Neita
status	sp. nov.

Treatment

Cyclocephala santandereana Neita , new species

( Figs. 2D View FIGURE 2 ; 15E View FIGURE 15 ; 16E View FIGURE 16 ; 17E View FIGURE 17 ; 18G View FIGURE 18 ; 24E View FIGURE 24 ; 25D View FIGURE 25 ; 27E–F View FIGURE 27 ; 39 View FIGURE 39 )

Type material. Holotype and paratype labeled “ Colombia, Santander, San Vicente / de Chucurí, Finca Familia Rincón , / 6º50´57.3´´N; 73º23´37.0´´W. WGS84 / 1214 m, Manual en Árbol de Cacao ( Theobroma cacao L.) / 29.iii.2018, D.F. Silva Tavera. IAvH-E-198644.” GoogleMaps Holotype deposited at IAVH; paratype deposited at UNSM .

Etymology. The species name “ santandereana ” refers to the type locality in Santander Province, Colombia and should be treated as an adjective in the nominative singular.

Diagnosis. Cyclocephala santandereana can be distinguished by the setose frons and clypeus ( Fig. 25D View FIGURE 25 ) (glabrous in C. brevipennis and C. fasciolata ( Fig. 25C View FIGURE 25 )); pygidium testaceous, with sparse punctures and slender, long setae ( Fig. 24E View FIGURE 24 ) (pygidium black in C. brevipennis and C. fasciolata ( Fig. 24D View FIGURE 24 )). The external edge at the base of protarsomere 5 with a small concavity ( Fig. 17E View FIGURE 17 ), lacking a small concavity in C. brevipennis and C. fasciolata ( Fig. 17C–D View FIGURE 17 ).

Description. Male holotype ( Fig. 2D View FIGURE 2 ). Length 21.4 mm; width 10.0 mm. Color black except each elytron and pronotum with transverse, slightly oblique band of reddish orange, band not reaching lateral or sutural margins; pygidium testaceous. Head: Frons and clypeus setose laterally ( Fig. 25D View FIGURE 25 ), with punctures moderate in size, becoming denser and smaller at clypeal apex. Clypeal apex broadly, weakly rounded, narrowly reflexed. Mandibles with concave inner edge, apex without teeth (as Fig. 9C View FIGURE 9 ); labium densely setose, paraglossa undeveloped, apex widely notched ( Fig. 12D View FIGURE 12 ); maxilla with galea developed, with 6 teeth (as Fig. 9D View FIGURE 9 ). Epipharynx, rectangular, densely setose, apex projecting at middle ( Fig. 14A–B View FIGURE 14 ). Antennal club subequal in length to antennomeres 2–7. Pronotum: Surface punctation similar to frons except punctures slightly larger. Base with strong marginal bead (Fig. 32A). Elytra: Surface with rows of moderately large, shallow punctures. Pygidium: Surface finely scabrous, dull, setigerous; setae moderately dense, long, tawny. In lateral view, surface regularly convex ( Fig. 24E View FIGURE 24 ). Legs: Protibia bidentate (18G). Protarsus enlarged ( Fig. 2D View FIGURE 2 ), tarsomeres 2–4 with large ventral lobes, all tarsomeres longitudinally striate on dorsal face, tarsomere 5 large, moderately curved ( Fig. 17E View FIGURE 17 ), without lobes or tooth, but lateral internal surface with longitudinal lines ( Fig. 16E View FIGURE 16 ), median claw large, strongly curved, apex finely cleft ( Fig. 15E View FIGURE 15 ), external edge with basal lines ( Fig. 17E View FIGURE 17 ). Length of metatarsus longer than metatibia. Venter: Prosternal process long, stout, apex obliquely flattened into a transverse oval with anterior 2/3 raised as convex “button”. Genitalia: Parameres as in Fig. 27E–F View FIGURE 27 .

Female. Unknown.

Paratype. Length 19.9 mm; width 9.9 mm. Similar to holotype except for the elytron and pronotum with transverse, slightly oblique band of reddish orange, band not reaching lateral or sutural margins, band occasionally broken in 4 spots or nearly obsolete

Distribution. Cyclocephala santandereana is known only from the type locality.

Life history. Adults are active during the day. The adults were taken on the leaf of Theobroma cacao L. ( Malvaceae ). The life history of this species is unknown.

Locality records ( Fig. 39 View FIGURE 39 ). 2 specimens examined with similar data. Colombia, Santander, San Vicente de Chucurí, Finca Familia Rincón .