Inocellia exusta, Cockerell & Custer, 1925, Cockerell & Custer, 1925

Makarkin, Vladimir N. & Archibald, S. Bruce, 2014, A revision of the late Eocene snakeflies (Raphidioptera) of the Florissant Formation, Colorado, with special reference to the wing venation of the Raphidiomorpha, Zootaxa 3784 (4), pp. 401-444 : 438-439

‘Fibla’ exusta ( Cockerell & Custer, 1925)

( Fig. 23 )

Inocellia exusta Cockerell & Custer, 1925: 296 , Fig.; Rodeck 1938: 290.

Fibla exusta: Carpenter 1936: 144 , 152, Fig. 13 ; Oswald 1990: 161; Engel 1998: 190; Engel 2003: 125; Aspöck & Aspöck 2004: 15, 17; Engel 2009: Tabl. 1; Engel 2011: 79 (Tabl. 2).

“ Inocellia ” exusta : Aspöck et al. 1991: 543, 666, Fig. 3060, Map 194.

“ Fibla ” exusta : Engel 2002: 17.

Redescription. Hind wing 8.6 mm long as preserved (estimated complete length ca. 9.5 mm), 3.1 mm wide. Costal space moderately broad; subcostal veinlets widely spaced. ScP relatively short; length from termination of ScP to pterostigma nearly equal to length of pterostigma. In subcostal space, one crossvein between ScP near its termination, RA. Pterostigma rather short. RA with one branch, closed pterostigma distally; posterior trace RA distad pterostigma angulated at 3ra-rp. Three crossveins between RA, RP; 2ra-rp located clearly proximad pterostigma; 3ra-rp short, distad pterostigma. Anterior trace of RP forked apically, with two branches proximad 3ra-rp. RP 1 deeply forked; RP 2 simple. One intraradial crossvein, between stem of RP 2, anterior branch of RP 1. Four crossveins between R, M: 1r-m very long, connecting RP, M at its separation from R; 2r-m connects stem of RP, M; 3r-m connects RP 1, MA; 4r-m connects posterior branch of RP 1, anterior branch of MA. M forked slightly proximad origin of RP. MA once deeply forked. MP pectinately branched; anterior trace simple; proximal-most branch (MP1) long, forked; two other branches short, simple. Three intramedian crossveins forming three doi; crossvein between anterior trace of MP, anterior branch of MP1 forming fourth doi. 1m-cu (‘M5’) long, oblique connecting basal MP, CuP; 2m-cu connects MP1 between origin, forking, and CuA. CuA pectinately forked, with two simple branches. CuP probably simple. Intercubital crossvein (icu) located proximad 1m-cu. AA1 with two simple branches.

Type material. Holotype UCM 4923 (coll. S.A. Rohwer). A well-preserved nearly complete hind wing. Verbatim label data [typed and hand written]: “F / B / UCM Paleontology TYPE Collection / Neuroptera : Raphidiidae / UCM #: 4923 / Locality #: 18130 Description: wing / ID: Inocellia exusta Cockerell / Age: Late Eocene Collector: SA Rohwer / Fm: Florissant Citation: Toohey”. The specimen has black ink writing on the rock that says “neurop. / UCM 4923a (AP) / R”; red ink writing “ TYPE ”, and a white hand written label glued to the rock that says “ Inocellia / exusta Ckll & Custer / TYPE [in red square] / Florissant / Miocene”.

Type locality. Florissant ( UCM locality No. 18130).

Remarks. The generic affinity of this species is provisional. The hind wing character states are fully concordant with those of the extant Fibla and Sininocellia , and the Baltic amber Succinofibla Aspöck et Aspöck, 2004 , in particular by the presence of the long basal r-m brace (1r-m), a plesiomorphic condition in Neuropterida. These genera are mainly distingiushed by genetalic features ( Aspöck & Aspöck 2004; Liu et al. 2012b).

Figures

FIGURE 13. Archiraphidia tranquilla (Scudder, 1890). Wing venation of the holotype MCZ 246. A, right forewing. B, right hind wing. C, left forewing. D, left hind wing. Scale bar = 4 mm (all to scale).

FIGURE 23. ‘ Fibla’ exusta (Cockerell et Custer, 1925). Holotype UCM 4923. A, specimen as preserved. B, hind wing venation. Scale bar = 4 mm.