Bengalia floccosa ( Wulp, 1885 )

Rognes, Knut, 2012, 3553, Zootaxa 3553, pp. 1-79 : 33-39

4. Bengalia floccosa ( Wulp, 1885)

Figs. 75–94.

Calliphora floccosa Wulp, 1885 : CCXCII. Two “female” syntypes from South Africa collected by de Selys-Fanson, and originally housed in IRSNB, are considered lost. Neotype male (BMSA, examined), here designated. Type locality: South Africa, KwaZulu-Natal, Ndumo Game Reserve . For details, see below under Type material.

Ochromyia crassirostris Karsch, 1888: 377 . Holotype female (MNHUB, examined), by monotypy [listed as “Usambara 1♀ ”; “type” or equivalent expression not used]. Type locality: Tanzania, Usambara.

Bengalia crassirostris: Bezzi 1908: 77 . Eritrea (doubtful record).

Note. Bezzi used this combination when citing Karsh’s paper (1888), even though Karsch did not (cf. previous entry).

Bengalia floccosa: Villeneuve 1914: 254 . Malawi, Zambia.

Note. Villeneuve described both males and females, remarking that “la coloration tire fréquemment sur le blond obscur”. Villeneuve cites material from “ Nyasaland ” (“Mt-Mlanje”, S.A. Neave), and “N. W. Rhodesia ”. He cites his own usage of Wulp’s name as “sec. type ”. What this expression exactly means is unclear. He may have compared the specimens before him with Wulp’s type(s), examined first hand, or compared them with Wulp’s description. No further details are given to explain its meaning. The interpretation of the “sec. typ.” expression that he actually examined Wulp’s types is perhaps too kind to him. If he did, he should have discovered and noted the mis-sexing of Wulp’s syntypes, see below under Type material.

Bengalia mercenaria Séguy, 1933: 78 . Holotype male (MNHN, examined), by original designation. Type locality: Mozambique (“Zambèze / Nova Choupanga / près Chemba”).

Bengalia floccosa: Zumpt 1956: 170 , 169 fig. 99. Reported from Democratic Republic of Congo, Tanzania, South Africa, Zimbabwe.

Bengalia floccosa: Zumpt 1956: 171 . One male and 1 female listed as type material from Usambara, but that is an error, only a single female type was described by Karsch from Usambara, see above under entry for O. crassirostris .

Bengalia floccosa: Pont 1980: 791 . Catalogue entry.

Afridigalia adrianponti Lehrer, 2005: 26 . Holotype male (BMNH, not examined), by original designation. Type locality: Tanzania (“Amani. Tanganyika”). Synonymy discussed in Rognes (2006).

Note. According to Lehrer’s description, A. adrianponti is based on a single dark specimen.

Afridigalia falsimonia Lehrer, 2005: 40 . Holotype male (BMNH, not examined), by original designation. Type locality: Malawi (as “ Nyasaland ”). Synonymy discussed in Rognes (2006).

Note. Afridigalia falsimonia is based on three pale specimens. Lehrer’s key (2005: 25) separates adrianponti from falsimonia on the shape of the antler as it appears in his drawings in lateral view. Obviously, in view of Lehrer’s (2011b) method of making his preparations (which involves heat, bergamot oil and Canada balsam), this fragile structure has assumed slightly different positions when Lehrer made the preparations, making chance events the basis for his species differentiation.

Afridigalia falsimonia: Lehrer 2006: 7 . South Africa, Angola, Democratic Republic of Congo, Kenya, Malawi (as “ Nyassaland ”), Tanzania, Zimbabwe.

Afridigalia zouloupyga Lehrer, 2006: 5 . Holotype male (BMNH, not examined), by original designation. Type locality: Malawi (as “ Nyassaland ”), Chitola Stream 10 mi W. of Domira Bay. Synonymy discussed in Rognes (2006).

Note. Afridigalia zouloupyga was based on two pale specimens with narrow dark posterior marginal bands on the abdominal tergites. No effort was made by Lehrer to describe the features by which it is supposed to differ from B. adrianponti or B. falsimonia . There are differences in the appearance of the antler in the published lateral-view drawings of B. zouloupyga , B. falsimonia and B. adrianponti , in addition to differences between the figures of the exact shape of the distal part of the ST5 flap, all of which I take to be result of individual variation. Interestingly, according to Lehrer (2006: 6) the holotype was captured by S.A. Neave and is labelled “ Bengalia floccosa Wulp , sec. typ.” in Villeneuve’s hand. It is thus a specimen Villeneuve seems to have identified on the basis of a comparison with the Wulp type (“sec. typ.”), although such a conclusion may be too kind to Villeneuve (see synonymy above, in note to the B. floccosa: Villeneuve, 1914 entry). Regarding the body colour of B. zouloupyga , B. falsimonia and B. adrianponti (pale or dark), Villeneuve (1914: 255) notes that whereas “ B. spinifemorata …et … B. Peuhi ” vary least in colour, other Bengalia species (including B. floccosa ) are “châtain, pouvant devenir tantôt pâlees avec une pruinosié clairsemée, tantôt foncées aved une pruinositée condensée [chestnut brown, sometimes pale with sparse pruinosity, sometimes dark with dense pruinosity]”. Nothing of this is discussed by Lehrer, who appears not have read Villeneuve’s papers.

Diagnosis. Male. Length: 10–13mm (n=5). Frons at vertex / head width ratio: 0.28–0.31 (mean 0.31, n=5). A species identifiable in the male sex by the unique shape and configuration of the upper lip and distal finger in the distiphallus, as seen in lateral view, in combination with the diagnostically shaped ST5 flap, but care should be taken to ensure that all of the posterolateral parts are observed.

Thorax often with dark pleura. Anepimeron with a bundle of dark setulae in upper part, lower part with numerous pale setulae. Femora usually dark, only one specimen seen has pale femora. Fore tibia with distinct spine-like setae ventrally in proximal half and a short fringe of thin setae distally. Mid tibia with a short fringe. Hind tibia with a dense fringe of long thin setae on the lower two thirds of av, v and pv surfaces.

Abdominal colour variable: pale with narrow dark bands, or very dark throughout, except that T1+2 and T3 shows slight yellowish translucence at middle on each side. A male in CNC has two strong and two weak discal setae on each side of the regular pair on T5.

ST5 flap with medial U-shaped excavation; the corner at the lateral edge of the excavation is rounded or sharp, whereas the corner where the remainder of the hind edge meets the lateral edge of the ST5 flap is evenly rounded.

Distal finger clearly visible in lateral view of the distiphallus, not hidden from lateral view by the vertical sheet of sclerotisation passing between the upper lip and base of antler; distal finger projecting forward and curving downward in lateral view, in dorsal view situated slightly laterad of the upper lip. Upper lip a square projection about as long as broad as seen from above and slightly curved in lateral view.

Female. Length: 8–12mm (n=4). Frons at vertex / head width ratio: 0.32–0.33 (mean 0.32, n=3). ST2 with pair of long strong black median marginal setae, a few weak setae outside each seta in pair, no black setae along lateral margin. Strong marginal pair of setae also on ST3 and ST4. Hind margin of T5 with an area of weak sclerotisation behind the marginal setae with an anterior delimitation shaped like a strongly curved edge, like in B. depressa (cf. Fig. 68). In ovipositor ST6 broad and undivided, as in B. depressa , but without a basal narrow “handle” set off from the rest of the ST6. ST7 with a large circular unsclerotised midventral area, occupying nearly the proximal two thirds of the length of ST7. Spermathecae relatively long and narrow compared to those of B. depressa . The cerci and epandrium short, not upturned, and with soft, not spinous, setae only [1 female dissected, a specimen collected together with males at the same time and place].

Discussion. The different appearance of the antler among Lehrer’s (2005) drawings of A. adrianponti , A. falsimonia and A. zouloupyga are likely to be artifacts, or due to different angles of view.

Biology. The material I have examined was collected from January–May, and from October–December. The dates given by Lehrer (2005, 2006) are within these ranges. Zumpt (1956) gives no dates outside these ranges. Thus B. floccosa seems not to be on the wing in the period from June to September. A male has been captured with an “hemerobe” ( Neuroptera , Hemerobiidae ) as its prey (holotype of B. mercenaria ). Nothing else is known about its biology and habits.

Distribution. Angola, Democratic Republic of Congo, * Kenya, Malawi, * Mozambique, * South Africa, * Tanzania, Zambia, * Zimbabwe. The records from Namibia by Kurahashi & Kirk-Spriggs (2006) are rejected because they were based on misidentified material; see below under B. tibiaria .

Material examined. Type material. Calliphora floccosa Wulp, 1885 . NEOTYPE male, here designated, in BMSA, labelled (1) Malaise traps / broad-leafed deciduous / woodland; (2) RSA: KZN, Ndumo Game R. / main camp area at: / 26°54.652'S 32°19.719'E / 27-30.xi.2009 / A.H.Kirk-Spriggs; (3) Entomology Dept. / National Museum / P. O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 13758; (5) NEOTYPE (m) / Calliphora floccosa / Wulp, 1885: CCXCII / K.Rognes des. 2012 [printed on red label]. The specimen has been dissected. The dried T1–5 are glued to a piece of card above the labels, the genitalia are in glycerol in a glass microvial above the neotype label.

Wulp based his species on “deux exemplaires femelles de l’Afrique australe (de Selys-Fanson)” (p. CCXCIII), de Selys-Fanson being the collector. He added that the specimens were “[p]armi les Diptères exotiques du Musée Royal d’Histoire Naturelle à Bruxelles [= IRSNB], que le zélé conservateur, M. Preudhomme de Borre , a bien voulu me confier dans le but de les examiner et de les déterminer,... ” (p. CCLXXXVIII). The syntypes are now not present in IRSNB, MRAC, RMNH or in ZMAN. They may not have been returned, and are probably lost .

There is good reason to assume that Wulp mis-sexed the specimens before him. He described the hind legs as having the “tibiis posticis hirsutis [hind tibia with long hairs]” (p. CCXCII), and “hérissées de longs poils au côté extérieur [with long hairs on the external side]”. Thus, in hindsight, we can say with confidence that he observed and reported the secondary male sexual characteristics of the hind legs. Being apparently unfamiliar with the genus Bengalia , he assigned his new species provisionally to the genus Calliphora . He noted that the elongate body, the total absence of metallic reflections from the abdomen and the peculiar pilosity of the hind legs might justify the erection of a separate genus for it. Most likely he thought his specimens were females because of the broad frons, “aussi large que les yeux [as broad as the eyes]”, since the males in Calliphora , unlike the males in Bengalia , usually have a very narrow frons.

Villeneuve possibly had seen one of the Wulp types; one of the specimens studied by Lehrer when erecting Afridigalia zouloupyga has a label to this effect. See entry for this nominal species in the synonymy, above.

No name-bearing types now exist for Calliphora floccosa Wulp, 1885 . The name has been in use for a long time, but has been involved in misidentifications, since never having been adequately characterised by reference to genital characters (one male in NMSA has been identified by Zumpt as B. gaillardi , another male from the same locality as B. cuthbertsoni , both identifications were made in 1979). It is involved in complex taxonomic problems involving several Afrotropical Bengalia species , i.e., the five species making up the B. floccosa subgroup of the B. peuhi species-group where, at present, not all females can be safely identified. Lehrer (2005: 18) has claimed that Calliphora floccosa is to be counted among nominal taxa “désignées, sans base scientifique, comme espèces valides”, because the description is based (allegedly) on unrecognisable females. Lehrer has obviously not read Wulp’s original description, and formed an independent opinion. Instead of fixing the interpretation of the old name by designating a neotype, or continuing the use of the name according to the understanding of the current authority ( Zumpt 1956), Lehrer (2005, 2006) described several new species under various names such as “ Afridigalia adrianponti ”, “ Afridigalia falsimonia ” and “ Afridigalia zouloupyga ”, even though all obviously belong to the same taxon as the one denoted by Zumpt (1956) as Bengalia floccosa (Wulp) on genital features. However, one should be aware that Zumpt himself (1956: 171) only distinguished male C. floccosa from other Bengalia males on the shape of the ST5 flap and the vestiture of the hind tibia. Even though he figured the distiphallus in profile view ( Zumpt 1956: 169 fig. 99 upper figure) in a way that clearly showed the unique configuration of the upper lip and distal finger, he never employed this feature to diagnose C. floccosa . In order to clarify the status of the nominal taxon C. floccosa I find it best to designate a neotype so that there no longer is any doubt about the identity of the name (Article 75.3.1 of ICZN 1999).

Wulp (1885: CCXCII–CCXCIII) described the abdomen as having “les deux premiers segments testacés à bord posterieur noir; les autres noirs, à reflets blanchâtres surtout à la moitié antérieurs des segments; ventre d’un brun rougeâtre [the two anterior segments testaceous with black hind margins; the others black, with whitish reflections particularly on the anterior half of the segments; ventrally reddish brown]”. The legs were described as being “fauves à cuisse noires; jambes postérieurs hérissées de longs poils au coté extérieurs [yellowish with black femora; hind legs densely clothed with long hairs on the external side]”. Thus it is evident that he had a rather dark specimen before him, with dark femora and abdomen with all dark T4 and T5 and partly dark T1+2 and T3. I have therefore selected a specimen from South Africa with these features as close to the description as I could find as neotype for Calliphora floccosa Wulp.

I here designate a specimen in BMSA (numbered 17758) as neotype for Calliphora floccosa Wulp, 1885 : CCXCII. It is labelled as cited above. Its characters fit the description, except that the T1+2 and T3 are very dark, only a small spot of yellow slips through the dark colouring. The type locality is in South Africa as were the specimens on which Wulp’s name was based. It has been dissected and the genitalia conform to the description above .

Ochromyia crassirostris Karsch, 1888: 377 . Karsch described O. crassirostris from a single female from Usambara (a mountain area in NE Tanzania). Holotype female, in MNHUB, labelled (1) Usambara / C. W. Schmidt / Febr – März 86 [printed on blue label]; (2) Type [printed on red label]; (3) 13841 [printed]; (4) Ochromyia / crassirostris / N. [handwritten; N. may mean Neu, or Nobis = our species]; (5) Bengalia ♀ / floccosa Wulp. / det. Zumpt 51 [Zumpt’s handwriting]; (6) Zool. Mus. / Berlin [printed]; (7) HOLOTYPE (f) / Ochromyia crassi - / rostris Karsch, 1888 / [“1887”]: 377 / K. Rognes 2012 [printed on red label]; (8) KR’s determination label as B. floccosa . ST2–4 each have a strong pair of median marginal setae as in Fig. 89. The specimen is of the pale type with yellow abdomen and thorax with narrow abdominal marginal bands.

Bengalia mercenaria Séguy, 1933: 78 . Holotype male, in MNHN, labelled: (1) Prédatrice avec / sa proie (Hémérobe) / 31 déc [handwritten on white label]; (2) COTYPE [red print on white label]; (3) MUSEUM PARIS / ZAMBÈZE / NOVA CHOUPANGA / PRÈS CHEMBA / P. LESNE 1929 [printed on blue label]; (4) Dissected by K. Rognes det. 2011 [printed label, except first line and year of last line which are handwritten]; (5) HOLOTYPE / Bengalia (m) / mercenaria / Séguy, 1933: 78 / (by original designation) / K. Rognes 19.iv.2011 [printed on red label]. Dissected by KR. Genitalia in glass microvial on pin below labels. Dried abdomen glued to card above labels, but below card bearing the prey’s wings. Paratype. MNHN: 1 male labelled (1) TYPE [red print on white label]; (2) Bengalia / mercenaria / Séguy / TYPE [handwritten by Séguy]; (3) PARATYPE / Bengalia (m) (not f!) / mercenaria / Séguy, 1933: 78 / (believed by Séguy / to be female) / K. Rognes 19.iv.2011 [printed on red label].

Séguy described B. mercenaria on the basis of specimens he believed to be of both sexes, but the two specimens under B. mercenaria in MNHN are both males. No symbols are present on the labels to show how Séguy sexed the specimens. Both specimens are in good condition with all legs intact. The male holotype of B. mercenaria is rather dark, and fits the description of Wulp of his syntypes of Calliphora floccosa . It is darker than the other, has a blue Lesne locality label and carries a pierced prey insect. This is clearly the one referred to by the words “… type ♂ (P. Lesne) capturé avec sa proie (Hémérobe)”. Séguy erroneously cites the date as “3 décembre”, whereas the uppermost handwritten label reads “31 déc.”. The loose wings of the “Hémérobe” were found in the box and were glued to a card below main part of the prey insect by KR. This specimen, even though labelled as COTYPE, is the holotype by original designation and I have labelled it as such. I have dissected it.

The second specimen, paler than the first, has no locality labels, but carries two Séguy “TYPE” labels. Even though a male I accept it as a paratype since it agrees with the description “ ♀: comme le mâle, de couleur plus pâle”. I have given it a red paratype label.

Other material. BMSA: South Africa: Eastern Cape: 1 female labelled (1) OOS – LONDEN [East London] / SE 3327 BB / 10 IV 1994 / H. VAN NUUREN [?] / DEPARTMENT OF ENTOMOLOGY / UNIVERSITY OF PRETORIA [handwritten, except last two lines which are printed]; (2) Ex Dept. of Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P. O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18156 [printed on white label, text facing down]; (5) KR’s determination label as B. floccosa . KwaZulu-Natal: 1 male 4 females labelled: (1) Malaise traps / broad-leafed deciduous / woodland: (2) RSA: KZN, Ndumo Game R. / main camp area at: / 26°54.652'S 32°19.719'E / 27-30.xi.2009 / A.H.Kirk-Spriggs; (3) Entomology Dept. / National Museum / P. O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 13757 / [1 male] / 13759 / 13760 / 13755 / 13756 (dissected) [4 females] [printed on white label, text facing down] (5) KR’s determination label as B. floccosa . The dissected female has the dried T1–5 glued to a card above the labels, the ovipositor and spermathecae are in glycerol in a glass microvial above label (5). 7 males 2 females labelled: (1) Malaise traps / sand & broadleafed / deciduous forest: (2) RSA: KZN, Ndumo Game R. / main road at: / 26°54.288'S 32°17.974'E / 4-8.xii.2009 / A.H.Kirk-Spriggs; (3) Entomology Dept. / National Museum / P. O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 16749 / 17032 / 17033 / 17034 / 17035 / 17152 / 17484 [7 males] / 17031 / 17153 [2 females] [printed on white label, text facing down]; (5) KR’s determination label as B. floccosa . 1 female labelled: (1) Malaise traps / grassy / floodplain; (2) RSA: KZN, Ndumo Game R. / pan at: / 26°54.288'S 32°17.974'E / 9-10.xii.2009 / A.H.Kirk-Spriggs; (3) Entomology Dept. / National Museum / P. O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 17483 [printed on white label, text facing down]; (5) KR’s determination label as B. floccosa . Limpopo: 1 male labelled (1) N-TVL [= Northern Transvaal] Tzaneen / RSA 6.iv.1996 / 30°10'S 23°50'E / Z. Liebenberg [printed on white label]; (2) Ex Dept. of Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P. O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18153 [printed on white label, text facing down]. Note. Pale specimen, femora hardly dark, narrow dark abdominal bands. Dissected by KR 7.i.2012.? province: 1 male labelled (1) SE 25 .. 31. Ac [? = South East 25 degrees 31 minutes of Ac] / 5-X-1980 / H.P. VAN WALT [?] / DEPARTMENT OF ENTOMOLOGY / UNIVERSITY OF PRETORIA [handwritten, except last two lines which are printed]; (2) Ex Dept. of Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P. O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18171 [printed on white label, text facing down]. Note. Pale specimen. Dissected by KR. Zimbabwe: 1 male labelled (1) P. MILES. / VUMBA / FEB. 1968 [handwritten, except first line which is printed in bluish letters]; (2) Ex Dept. of Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P. O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 11353. Note. Pale specimen. CNC: Tanzania: 1 male labelled (1) TANGANYIKA / Same / 25 [handwritten] Mai 1962; (2) KR’s determination label; 1 male labelled (1) TANGANYIKA / Same / Mai 1962; (2) KR’s determination label. IRD: Zimbabwe: 1 male labelled (1) Chikwedziwa / Rhodesie / HBS rec 10/59 [handwritten]; (2) KR’s determination label. Measurements: Frons at vertex / head width ratio: 39 / 120, length: 11mm.? country: 1 male labelled (1) KR’s determination label. [No locality label]. Measurements: Frons at vertex / head width ratio: 37 / 120, length: 11mm. KR: Kenya: Coast Province: 1 male labelled (1) KENYA – Coast province / 7 KM W Kakoneni, 100 m / UTM 37 M 588225 9647804 / 24.V.2006 (hand net) / D.Avesani, G.Carpaneto, G.Nardi / & P.Cerretti leg. [printed]; (2) KR’s determination label as B. floccosa . Dissected by KR. 1 male labelled (1) Kenya – Coast province / N edge of Arabuko Sokoke Forest / UTM 37 M 607257 9644873 / 83 m, 24.V.2006 (hand net) / D.Avesani, G.Carpaneto, G.Nardi / & P.Cerretti leg. [printed]; (2) KR’s determination label as B. floccosa . MNHUB: Tanzania (Iringa Region): 1 male labelled (1) Nyassa-See / Langenburg / 3.X.-4.XI.96 / Fülleborn S [blue printed label; the year is 1896]; (2) crassirostris [handwritten label]; (3) Bengalia ♂ / floccosa Wulp. / det. Zumpt 51 [handwritten by Zumpt]; (4) Zool. Mus. / Berlin [printed]; (5) My determination label as B. floccosa . Note. The current name for Langenburg is Lumbira at Lake Nyasa, Iringa Region, Tanzania. This specimen is of the pale type with yellow abdomen and thorax with narrow abdominal marginal bands. NMSA: South Africa: KwaZulu-Natal: 1 male labelled (1) South Africa, Natal Prov / Zululand, 20 mi. S.Ndumu / Game Res.Camp (2732Aa) / Nov.29.1971 ( ME &BJ Irwin / dry scrub forest; 320ft. [printed white label]; (2) Bengalia ♂ / gaillardi S & G. / det Zumpt 79 [Zumpt’s handwriting]; (3) NMSA- DIP / 17708 [white label, print facing down]; (4) NATAL MUSEUM / Pietermaritzburg / South Africa [printed on green label]; (5) KR’s determination label as B. floccosa . Note. Zumpt had extracted the epandrium with cerci and surstyli and glued the complex to a card above the labels. The ST5 flap was intact and clearly visible on the tip of the abdomen when I received the specimen. T1–5 very dark, with only traces of yellow shining through on T1+2 and T3 laterally. I have dissected it. ST1–5 and the remainder of the genitalia in glycerol in glass microvial on pin above label (5). Dried T1–5 glued to same card as the extracted cerci and surstyli. 1 male labelled (1) South Africa: Natal / Ndumu Game Reserve / Rest camp 2632Cd 95m / 23–9:XI:1977 Malaise / Brothers& J.Guillardmod [printed]; (2) Bengalia ♂ / cuthbertsoni Zpt. / det Zumpt 79 [handwritten by Zumpt]; (3) NMSA- DIP / 17843 [white label, print facing down]; (4) NATAL MUSEUM / Pietermaritzburg / South Africa [printed on green label]; (5) KR’s determination label as B. floccosa . Note. The cerci and surstyli have been extracted by Zumpt and are glued to a card above the labels, the left surstylus is absent. The ST5 flap is intact and clearly visible at the tip of the abdomen. Aedeagus appears to be intact inside the tip of the abdomen, but not extracted by Zumpt. Abdomen and femora all dark. Head glued to thorax and there is a lot of glue on right fore leg. The spine-like setae on the ventral side of the right fore tibia visible through the glue. Left fore leg, right mid leg and left hind tibia lost. I have dissected the abdomen. ST1–5 and the remainder of the genitalia in glycerol in glass microvial on pin above label (5). Dried T1–5 glued to same card as the extracted cerci and surstyli. 1 female labelled (1) Tete Pan // Zululand // leg. Paterson [printed on upper side] 26/4/55 [pencil writing on underside]; (2) Bengalia ♀ / floccosa Wulp. / det. Zumpt 55 [handwritten by Zumpt]; (3) NMSA-DIP / 17849 [printed, face down]; (4) NATAL MUSEUM / Pietermaritzburg / South Africa [printed on green label]; (5) KR’s determination label. Note. The specimen was placed in collection under B. depressa . It has a pair of strong black setae along hind margin of ST2 (opposite the condition in B. depressa ), and a conspicuous area of weak sclerotisation at the hind margin of T5 (as in B. depressa ). RMNH / ZMAN: Kenya: 1 male labelled (1) MALINDI / 5 & 6. I. 1968 [printed]; (2) KENYA / Kilifi / D. Gillissen / L. Blommers; (3) KR’s determination label. Dissected by KR. Genitalia and abdominal sternites in glass microvial in glycerol above label (3). Dried abdominal tergites glued to card above label (1). A mid leg also glued to the card.