Blennidus
publication ID |
https://doi.org/ 10.11646/zootaxa.4000.1.1 |
publication LSID |
lsid:zoobank.org:pub:00769C76-A2D4-4792-A5DD-5F2B7EA352CB |
DOI |
https://doi.org/10.5281/zenodo.6096114 |
persistent identifier |
https://treatment.plazi.org/id/039087C7-FFBB-FF90-E28C-49A030B56A69 |
treatment provided by |
Plazi |
scientific name |
Blennidus |
status |
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Key to the species of Blennidus View in CoL subgenus Agraphoderus recorded from Peru
(To cope with a possible misinterpretation at couplet 27, Blennidus anxius is keyed twice)
1. Pronotum flattened on disc and markedly restricted at base, with sides rectilinear or slightly sinuate in posterior third; hind angles of pronotum obtuse, with blunt apex ( Fig. 62 View FIGURES 58 – 65 ). Elytra subdepressed. Aedeagus curved in dorsal view and with apical blade sharply pointed ( Fig. 98 View FIGURES 94 – 99 ). Dept. Ancash, Cordillera blanca. 8.3–10.3 mm ...... Blennidus huascarani Allegro, 2010 View in CoL .
- Species dorsally convex. The above characters are not all present together....................................... 2.
2. Male mesotibiae with an inner spine or an acutely pointed swelling in subapical position........................... 3.
- Male mesotibie without an inner subapical spine, at most with an obtuse swelling................................. 5.
3. Male metatibiae with a marked subapical swelling. Hind angles of pronotum widely rounded (Fig. 54). Male dorsally with strong metallic lustre. Dept. Cajamarca. 9.5–9.8 mm .............................. Blennidus bellesi ( Straneo, 1993) View in CoL
- Male metatibiae subapically not swollen. Hind angles of pronotum obtuse but more or less indicated.................. 4.
4. Hind angles of pronotum with pointed vertex ( Fig. 68 View FIGURES 66 – 73 ). Elytra subparallel sided or slightly dilated backwards, with rounded apex. The subapical inner spine of male mesotibiae short. Aedeagus short and markedly curved downwards at apex in lateral view ( Fig. 104 View FIGURES 100 – 105 ). Dept. Cuzco. 8.5–9.5 mm .................................. Blennidus mesotibialis ( Straneo, 1993) View in CoL
- Hind angles of pronotum with blunt vertex ( Fig. 69 View FIGURES 66 – 73 ). Elytra oval, with pointed apex. The subapical inner spine of male mesotibia long and acuminate. Aedeagus slender, scarcely curved downwards at apex in lateral view ( Fig. 105 View FIGURES 100 – 105 ). Dept. Cajamarca. 9.5 mm ............................................................... Blennidus negrei ( Straneo, 1993) View in CoL
5. Pronotum usually disc-shaped, with hind angles more or less rounded (Figs. 56, 72, 76, 78). Species distributed in northern Peru (Dept. Cajamarca)............................................................................... 6.
- Pronotum of different shape, or species distributed in central or southern Peru.................................... 9.
6. The 1st to 5th elytral striae all equally impressed, with moderately convex intervals. Hind angles of pronotum rounded or hardly evident ( Fig. 76 View FIGURES 74 – 81 ). Larger size (9.2–10.9 mm).......................... Blennidus pinguis (Allegro & Giachino, 2011) View in CoL
- The 1st elytral stria more impressed than 2nd to 5th, which are more or less obsolete, with flat intervals. Hind angles of pronotum rounded or widely obtuse. Smaller size (<9 mm)........................................................... 7.
7. Pronotum with base as wide as anterior margin, with sides strongly convex and hind angles rounded (Fig. 56). 9.0 mm. Dept. Cajamarca............................................................... Blennidus crassus ( Straneo, 1993) View in CoL
- Pronotum with base definitely narrower than anterior margin, with sides less convex and hind angles widely obtuse but evident ( Figs. 72 View FIGURES 66 – 73 , 78 View FIGURES 74 – 81 )........................................................................................ 8.
8. Larger size (8.5–9.0 mm). Umbilicate pores at sides of elytra widely interrupted at middle. Sides of pronotum linear or slightly convex just before hind angles ( Fig. 72 View FIGURES 66 – 73 ). Sterna IV–VI with a series of evident punctures at sides. Aedeagus in dorsal view markedly arcuate ( Fig. 108 View FIGURES 106 – 111 )................................................ Blennidus orbicollis ( Straneo, 1993) View in CoL
- Smaller size (7.5–8.0 mm). Umbilicate pores at sides of elytra not widely interrupted at middle. Sides of pronotum slightly sinuate just before hind angles ( Fig. 78 View FIGURES 74 – 81 ). Sterna IV–VI without punctures at sides. Aedeagus in dorsal view moderately arcuate ( Fig. 114 View FIGURES 112 – 117 ).................................................. Blennidus pseudolaevis (Allegro & Giachino, 2011) View in CoL
9. Male fore tarsi scarcely dilated, without scaly pubescence underneath. Head very large. Pronotum clearly transverse (W/ L>1.4) ( Fig. 63 View FIGURES 58 – 65 ). The 1st elytral stria definitely more impressed than others. Large body size (11–13 mm). Dept. Pasco (Cerro de Pasco)............................................................ Blennidus inca ( Tschitschérine, 1898) View in CoL
-.. Male fore-tarsi normally dilated, with scaly pubescence underneath. The other above characters not all present together 10.
10. Pronotum widely flattened before hind angles, which are angulately obtuse and clearly raised ( Fig. 80 View FIGURES 74 – 81 ). Elytra with subparallel sides and striae all equally impressed. 9.5 mm. Unknown distribution............ Blennidus refleximargo ( Straneo, 1993) View in CoL
- Pronotum with different shape......................................................................... 11.
11. Elytra with a single discal setigerous pore on 3rd interval and with marked green or bluish metallic lustre.............. 12.
- Elytra with at least two discal setigerous pores on 3rd interval. Brown or black species, at most with slight metallic lustre. 14.
12. Elytra with green-bluish lustre. Striae superficially impressed but evident; the 1st, as well as the 8th and 9th, more impressed than others. Sides of pronotum regularly rounded with a very shortly sinuation before basal angles, which are obtusely angulate ( Fig. 84 View FIGURES 82 – 87 ). 9 mm. Unknown distribution.................................... Blennidus tenenbaumi ( Lutshnik, 1927) View in CoL
- Elytra with marked green-olive lustre................................................................... 13.
13. Pronotum with basal angles right and acutely pointed ( Fig. 71 View FIGURES 66 – 73 ). Head with frontal furrows very faint. Elytral striae all very superficial. 9 mm. Unknown distribution............................... Blennidus olivaceus ( Tschitschérine, 1897) View in CoL
- Pronotum with basal angles obtuse, with a small blunt tooth at vertex. Head with frontal furrows impressed. The 1st elytral stria, as well the 7th–9th, more impressed than others. 9 mm. Unknown distribution.................................................................................................... Blennidus idioderus ( Tschitschérine, 1898) View in CoL
14. Lateral margins of pronotum with two close setigerous pores on anterior half. Posterior angles of pronotum rounded ( Fig. 60 View FIGURES 58 – 65 ) 8.5 mm. Unknown distribution...................................... Blennidus euphaenops ( Tschitschérine, 1898) View in CoL
- Lateral margins of pronotum with a single setigerous pore on anterior half. Posterior angles of pronotum rounded or angulate.................................................................................................. 15.
15. Species ranging in the mountains of Cordillera Blanca (Dept. Ancash). Only the 1st elytral stria well impressed, the others superficial or obliterated. Male mesotibiae preapically swollen. Seta on posterior angles of pronotum not moved forewards.................................................................................................... 16.
- Species recorded from other Andean areas of Peru. The above characters not all present together.................... 17.
16. Posterior angles of pronotum almost rounded ( Fig. 86 View FIGURES 82 – 87 ). Larger size (8.5–9.4 mm). Median lobe of aedeagus stouter ( Fig. 119 View FIGURES 118 – 120 )........................................................................ Blennidus unistria ( Straneo, 1993) View in CoL
- Posterior angles of pronotum obtuse but clearly indicated by a small tooth ( Fig. 61 View FIGURES 58 – 65 ). Smaller size (7.4–9.2 mm). Median lobe of aedeagus slender ( Fig. 97 View FIGURES 94 – 99 ).................................... Blennidus fitzcarraldi (Giachino & Allegro, 2011) View in CoL
17. Species ranging in the Andes of southern Peru. Elytral striae all equally impressed (at most the 1st slightly more impressed). Male mesotibiae not preapically swollen (sometimes metatibiae preapically swollen). Seta on posterior angles of pronotum not moved forewards................................................................................... 18.
- Species recorded from other Andean areas of Peru. The above characters not all present together.................... 23.
18. Sides of pronotum sinuate at the base or posteriorly nearly straight; male metatibiae not preapically swollen........... 19.
- Sides of pronotum rounded over entire length; male metatibiae preapically swollen or not.......................... 22.
19. Impressed elytral striae with feebly convex intervals in ♂. Distinctly pear-shaped elytra. Aedeagus with median lobe regularly curved and not depressed ventrally ( Fig. 100 View FIGURES 100 – 105 ). Species from the Puno area. 7.7–7.8 mm ................................................................................................... Blennidus jelskii ( Tschitschérine, 1897) View in CoL
- Superficial elytral striae with flat intervals in both sexes. Elytra not pear-shaped. Aedeagus with median lobe angulately inserted on the basal bulb and ventrally somewhat depressed. Species from the Cuzco area........................ 20.
20. Sides of pronotum markedly sinuate at base (Fig. 51). A faint metallic lustre is usually present in ♂. In lateral view, median lobe of aedeagus larger and distally nearly straight ( Fig. 89 View FIGURES 88 – 93 ). Relatively larger (8.0–9.0 mm)..................................................................................... Blennidus abramalagae (Allegro & Giachino, 2011) View in CoL
- Sides of pronotum weakly sinuate or nearly straight at base. Metallic lustre usually absent. Median lobe of aedeagus smaller and angulately bent downwards in the distal part ( Figs. 113, 117 View FIGURES 112 – 117 ). Relatively smaller (6.6–8.8 mm).................. 21.
21. Habitus more slender (length/width of elytra = 1.50–1.57). Elytra less convex, with apical declivity less marked. Aedeagus smaller even in the largest specimens ( Fig. 113 View FIGURES 112 – 117 ). 6.6–8.8 mm ............ Blennidus procerus (Allegro & Giachino, 2011) View in CoL
- Habitus oval (length/width of elytra = 1.45). Elytra more convex, with marked apical declivity. Aedeagus more slender and larger ( Fig. 117 View FIGURES 112 – 117 ). 7.7–8.7 mm ...................................... Blennidus straneoi (Allegro & Giachino, 2011) View in CoL
22. Elytral striae definitely impressed, with slightly convex intervals. Male metatibiae preapically not swollen. Median lobe of aedeagus distally rounded in dorsal view ( Fig. 91 View FIGURES 88 – 93 ). Larger size (9.5 mm). Species from the Puno area.................................................................................. Blennidus aulacostigma ( Tschitschérine, 1897) View in CoL
- Elytral striae superficial, with flat intervals. Male metatibiae preapically swollen. Median lobe of aedeagus distally pointed in dorsal view ( Fig. 95 View FIGURES 94 – 99 ). Smaller size (7.6–9.3 mm). Species from the Cuzco area.................................................................................................... Blennidus etontii (Allegro & Giachino, 2011) View in CoL
23. Hind angles of pronotum more or less rounded, always blunt at apex, and postangular seta distinctly moved forewards... 24.
- Hind angles of pronotum rounded or angulate; if rounded the postangular seta not moved forewards................. 27.
24. The 1st elytral stria distinctly more impressed than others; intervals flat. Elytra oval shaped, narrow at base and dilated backwards; shoulders rounded. Male mesotibiae not preapically swollen. The apical blade of aedeagus slightly bent to left in dorsal view ( Fig. 118 View FIGURES 118 – 120 ). Ticlio, Abra de Anticona..................................... Blennidus ticlianus ( Straneo, 1993) View in CoL
- Elytral striae well impressed and intervals subconvex; the 1st stria only slightly more impressed than others. Male mesotibiae preapically swollen. The apical blade of aedeagus not laterally bent in dorsal view. Species from Depts. Pasco and Huanuco.................................................................................................... 25.
25. Male metatibiae with a marked subapical swelling. Depts. Pasco and Huanuco.. Blennidus languens ( Tschitschérine, 1898) View in CoL
- Male metatibiae without marked subapical swelling........................................................ 26.
26. Larger size (10–12 mm). Elytra oval shaped, narrower at base and dilated backwards. Front angles of pronotum not prominent ( Fig. 74 View FIGURES 74 – 81 ).............................................................. Blennidus pascoensis ( Straneo, 1954) View in CoL
- Smaller size (9–9.5 mm). Elytra shorter, with subparallel sides, not distinctly dilated backwards. Front angles of pronotum prominent (Fig. 55)........................................................... Blennidus bombonensis View in CoL n. sp.
27. Hind angles of pronotum broadly or narrowly rounded...................................................... 28.
- Hind angles of pronotum from almost right (sometimes blunt at vertex) to obtuse, but always angulate................ 34.
28. Larger size (> 11 mm). Elytral striae more or less equally impressed. Posterior angles of pronotum broadly rounded ( Fig. 67 View FIGURES 66 – 73 ).
Depts. Junin and Huancavelica............................................... Blennidus mateui ( Straneo, 1993) View in CoL - Smaller size (<10.5 mm). Posterior angles of pronotum more or less narrowly rounded. The 1st elytral stria usually more impressed than others; intervals flat..................................................................... 29.
29. Elytra short-oval, relatively wide (L/W=1.40). Sides of pronotum arcuate and hind angles broadly obtuse ( Fig. 82 View FIGURES 82 – 87 ). 7.1 mm. Dept. La Libertad........................................................ Blennidus sublaevis ( Straneo, 1993) View in CoL
- Elytra more slender (L/W=1.50). Body larger (> 7.5 mm).................................................... 30.
30. Smaller (7.5–8.2 mm). Apical blade of aedeagus short and almost triangular in dorsal view......................... 31.
- Larger (9.1–10.2 mm). Apical blade of aedeagus longer, widely truncate at apex and bent to the right in dorsal view ( Figs. 88 View FIGURES 88 – 93 , 102 View FIGURES 100 – 105 ).............................................................................................. 33.
31. Body dark brown. Legs reddish-brown. Elytral striae 2–5 generally superficial, distinctly less impressed than 1st. Dept. Junin..................................................................... Blennidus vereshaginae ( Straneo, 1993) View in CoL
- Body piceous black, with concolorous legs. Striae 2–5 complete and distinct, although less impressed than 1st.......... 32.
32. Posterior angles of pronotum rounded ( Fig. 83 View FIGURES 82 – 87 ). Unknown distribution........... Blennidus tardus ( Tschitschérine, 1898) View in CoL
- Posterior angles of pronotum obtuse with blunt vertex (Fig. 52). Dept. Lima...... Blennidus anxius ( Tschitschérine, 1898) View in CoL
33. Males usually with bronze lustre. Aedeagus not exceptionally slender ( Fig. 88 View FIGURES 88 – 93 ). Dept. Huancavelica................................................................................... Blennidus abditus (Giachino & Allegro, 2011) View in CoL
- Males usually without bronze lustre. Aedeagus exceptionally long and slender, very distinctive ( Fig. 102 View FIGURES 100 – 105 ). Dept. Junin............................................................................. Blennidus longiloba ( Straneo, 1993) View in CoL
34. Sides of pronotum markedly convex, widest at anterior third ( Figs. 58 View FIGURES 58 – 65 , 73 View FIGURES 66 – 73 ). The 1st elytral stria definitely more impressed than others, which are very superficial; intervals flat........................................................... 35
- Sides of pronotum moderately convex, sometimes linear towards base, widest at middle or just beyond middle. The 1st elytral stria hardly more impressed than others, which are complete and distinct; intervals flat or subconvex................. 36.
35. Smaller size (7 mm). Sides of pronotum less rounded in basal half, with very short sinuation at base ( Fig. 58 View FIGURES 58 – 65 ). Unknown distribution.............................................................. Blennidus egens ( Tschitschérine, 1898) View in CoL
- Larger size (8.4 mm). Sides of pronotum evenly rounded on whole length ( Fig. 73 View FIGURES 66 – 73 ). Dept. Junin........................................................................................ Blennidus pachycerus ( Tschitschérine, 1897) View in CoL
36. Pronotum subquadrangular with sides hardly convex, almost linear or subsinuate before hind angles (Figs. 57, 70, 75)... 37.
- Pronotum subquadrangular or transverse, with sides more convex and usually evenly rounded before hind angles....... 39.
37. Elytra short-oval, dull even in males due to a strong microsculpture; striae distinctly punctate and very superficial in posterior third. 8–9 mm. Depts. Pasco and Junin......................................... Blennidus curtatus ( Straneo, 1993) View in CoL
- Elytra oval-elongate, moderately shiny in males; striae impunctate and well impressed even in posterior third.......... 38.
38. Larger size (8.5–9 mm). Sides of pronotum linear (or slightly convex) in basal half ( Fig. 70 View FIGURES 66 – 73 ). Elytra angulate at shoulders. Median lobe of aedeagus shorter and stouter ( Fig. 106 View FIGURES 106 – 111 ). Depts. Junin-Pasco......... Blennidus nigritulus ( Straneo, 1993) View in CoL
- Smaller size (7.6 mm). Sides of pronotum with short sinuation before hind angles ( Fig. 75 View FIGURES 74 – 81 ). Elytra rounded at shoulders. Median lobe of aedeagus more slender ( Fig. 111 View FIGURES 106 – 111 ). Unknown distribution.... Blennidus phaenogonus ( Tschitschérine, 1898) View in CoL
39. Smaller size (8.2 mm). Fore angles of pronotum completely obliterated (Fig. 52). Median lobe of aedeagus hardly curved downwards at apex in lateral view ( Fig. 90 View FIGURES 88 – 93 ). Dept. Lima...................... Blennidus anxius ( Tschitschérine, 1898) View in CoL
- Larger size (8.5–9 mm). Fore angles of pronotum more or less prominent ( Fig. 79 View FIGURES 74 – 81 ). Median lobe of aedeagus distinctly curved downwards at apex in lateral view ( Fig. 115 View FIGURES 112 – 117 ). Dept. Junin...................... Blennidus rectangulus ( Straneo, 1993) View in CoL
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Pterostichinae |
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Pterostichinae |
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