Eutyphlus LeConte, 1880

Owens, Brittany E. & Carlton, Christopher E., 2016, Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini, The Coleopterists Bulletin 70 (1), pp. 1-29 : 1-29

Eutyphlus LeConte, 1880

( Figs. 1, 2 )

Redescription. Possessing characters of the tribe Trichonychini (sensu Reitter 1882) ( Figs. 1, 2 ). MALE. Integument: Body and appendages uniformly light brown, evenly covered by sparse, short, suberect pubescence. Capitate genal setae present. Head: Vertexal foveae present, large, setose, located posterior to shallow, U-shaped vertexal sulcus. Antennal acetabula remote, antennomeres 1–10 unmodified, antennomere 11 with lateral, shelf-like excavation bearing dense cluster of setae. Labrum wider than long, slightly expanded distally, apical margin gently concave, bearing

Character Number

00000000011111111112222222222333333333344444444445555 Taxa 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 Arthmius bulbifer 1 0 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Hendecameros panamense 1 0 0 0 0 1 0 0 0 0 0 1 2 1 0 3 1 0 0 1 0 1 1 1 1 1 1 2 2 0 1 1 1 1 1 1 1 1 1 0 0 0 1 0 0 2 0 0 1 1 0 0 1 Barroeuplectoides sp. 1 1 0 0 0 0 0 1 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 1 0 0 1 1 0 0 0 0 0 1 Euplectus confluens 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 2 1 0 0 0 0 1 0 0 1 0 0 1 1 0 0 1 Leptoplectus pertenuis 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 1 1 1 0 1 0 1 2 1 0 0 0 0 1 0 0 1 0 0 0 1 0 0 1 Pycnoplectus linearis 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 2 1 0 0 0 0 1 0 0 1 0 0 1 1 0 0 1 Jubus sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 2 1 1 2 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 1 0 0 0 0 0 1 1 1 0 1 1 0 0 1 Sebaga sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 2 1 1 2 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 1 0 0 0 0 0 1 1 0 0 1 1 0 1 1 Mayetia domestica 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 1 0 3 1 1 1 1 1 2 2 1 1 1 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 1 Metopias sp. 1 0 0 0 0 1 1 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 1 1 2 1 0 0 0 0 0 0 0 1 0 0 0 0 0 0 1 Foveoscapha terracola 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 1 0 1 1 0 0 1 1 0 1 1 Bibloplectus cherokee 1 1 0 3 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 0 1 0 0 0 1 0 0 1 Eutyphlus dybasi 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 0 1 1 0 1 Eutyphlus spiralis 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 0 1 1 0 1 Eutyphlus prominens 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 1 0 1 0 0 0 1 1 0 1 Eutyphlus schmitti 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 1 0 1 0 0 0 1 1 0 1 Eutyphlus similis 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 0 1 1 0 1 Eutyphlus thoracicus 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 0 1 0 0 0 1 1 0 1 Pseudactium carolinae 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 1 1 0 0 1 Rhinoscepsis bistriatus 1 1 0 0 0 1 1 0 0 0 0 0 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 1 1 0 0 1 0 1 1 1 0 1 1 0 0 1 Thesiastes sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 1 0 0 1 0 0 1 1 0 0 1 Thesium sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 1 1 0 0 1 0 1 1 0 0 1 1 0 0 1 Trimioplectus australis 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 1 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 1 0 0 0 0 1 0 1 1 Actiastes globiferum 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 0 1 0 0 1 1 1 1 1 Dalmosanus quercavum 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 1 0 0 0 0 1 0 0 1 Dalmosella tenuis 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 0 1 0 0 2 1 0 0 1 Hanfordia sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 1 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 Melba clypeata 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 1 0 0 0 Trimiomelba dubia 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 Trimium brevicorne 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 1 0 0 0 Quadrelba sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 1 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 Conoplectus acornus 1 1 0 0 0 0 0 0 0 2 0 1 0 1 0 1 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 0 1 0 1 1 Rhexius sp. 1 1 0 0 0 0 0 0 0 2 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 2 1 0 1 1 1 0 2 1 0 1 1 Sonoma baylessae 1 0 1 3 1 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 0 1 1 1 1 1 1 1 2 0 1 1 0 1 3 1 1 1 2 2 0 0 0 0 0 0 0 0 0 0 0 0 1 Reichenbachia howardi 1 0 0 3 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 2 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Pselaptrichus perfidus 1 1 0 0 0 1 1 0 1 0 1 1 0 1 0 0 1 0 0 1 0 2 4 1 4 1 1 1 3 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 Lucifotychus cognatus 1 1 0 0 0 1 1 0 0 0 0 1 0 1 0 0 1 0 0 2 0 2 1 2 1 2 3 2 3 1 1 1 0 1 0 1 2 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 1

Character Number

55555566666666667777777777888888888899999999991111111

Taxa 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 Arthmius bulbifer 0 0 1 1 1 0 0 1 0 1 1 1 1 0 0 0 0 0 1 0 0 0 1 1 0 1 0 0 0 0 1 1 0 1 0 0 0 1 1 1 0 0 0 0 0 3 0 0 0 0 0 1 1 Hendecameros panamense 0 0 1 0 1 0 1 1 0 1 1 1 1 1 0 0 1 0 0 0 1 1 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 0 1 0 0 0 0 0 0 1 0 0 1 0 0 2 1 Barroeuplectoides sp. 1 0 0 1 1 0 1 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 1 0 0 1 0 0 0 0 0 0 0 4 0 0 1 0 0 2 1 Euplectus confluens 1 0 0 2 1 0 0 1 0 0 1 1 1 1 0 1 0 0 1 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 1 1 0 1 0 1 4 0 0 0 0 0 1 1 Leptoplectus pertenuis 1 0 0 3 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 1 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 4 0 0 0 0 0 1 1 Pycnoplectus linearis 0 0 0 1 1 0 0 1 0 0 1 1 1 1 0 1 0 0 1 0 0 0 1 1 0 0 0 0 1 0 0 1 0 0 0 0 0 0 1 1 0 0 1 1 1 4 0 0 0 0 0 1 1 Jubus sp. 0 0 0 3 2 0 0 1 0 0 1 1 1 1 0 0 1 0 0 0 0 0 1 0 0 0 0 0 1 1 1 2 0 0 1 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Sebaga sp. 0 0 0 3 2 0 0 1 0 0 1 1 2 1 0 0 1 0 0 0 0 0 1 0 0 0 0 0 1 1 1 2 0 0 1 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Mayetia domestica 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1 1 0 0 0 0 0 1 1 0 2 0 1 0 0 0 0 0 0 0 0 0 0 0 4 0 0 1 0 0 2 1 Metopias sp. 0 0 0 0 1 0 0 1 1 1 1 1 1 0 0 1 0 0 0 0 1 0 1 1 1 0 0 0 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 Foveoscapha terracola 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 1 1 Bibloplectus cherokee 0 0 0 0 1 0 0 1 0 1 1 1 1 1 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 1 0 0 0 0 0 1 0 2 0 0 0 0 0 0 0 0 0 0 0 1 1 Eutyphlus dybasi 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 1 1 1 0 0 0 1 1 1 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Eutyphlus spiralis 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 1 1 1 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Eutyphlus prominens 1 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 1 1 1 1 0 1 3 0 0 0 0 0 1 1 Eutyphlus schmitti 1 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 1 1 0 1 1 1 3 0 0 0 0 0 1 1 Eutyphlus similis 1 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 1 0 0 0 0 0 3 0 0 0 0 0 1 1 Eutyphlus thoracicus 1 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 1 0 0 0 0 0 3 0 0 0 0 0 1 1 Pseudactium carolinae 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 1 0 1 0 0 0 1 1 1 0 0 0 1 1 1 2 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Rhinoscepsis bistriatus 0 0 0 0 1 0 0 1 1 1 1 1 2 1 0 2 0 0 1 0 0 0 1 1 0 0 0 0 1 1 1 2 0 0 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Thesiastes sp. 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 1 0 1 0 0 0 1 1 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0 1 0 0 1 0 1 0 0 0 0 0 0 1 1 Thesium sp. 1 0 0 3 1 0 1 1 0 1 1 1 1 1 0 1 1 2 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Trimioplectus australis 0 0 0 2 2 0 0 1 0 0 1 1 2 0 0 1 1 0 1 0 0 0 1 1 0 1 1 0 0 0 1 1 0 0 0 0 0 1 0 0 1 1 1 1 0 0 0 0 0 0 0 1 1 Actiastes globiferum 0 0 0 1 1 0 0 1 0 0 1 1 1 1 0 1 1 0 1 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 2 0 0 0 0 0 0 0 0 0 0 0 1 1 Dalmosanus quercavum 0 0 0 3 1 0 0 1 0 1 1 1 2 1 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 1 1 1 0 1 0 0 0 1 0 2 0 0 0 0 1 0 0 0 0 0 0 1 1 Dalmosella tenuis 0 0 0 0 1 0 0 1 0 0 1 1 2 1 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 2 0 0 0 0 0 0 0 0 0 0 0 1 1 Hanfordia sp. 0 0 0 3 2 0 0 0 0 0 1 1 2 0 0 0 0 0 1 0 0 0 1 1 1 1 0 0 1 1 1 2 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Melba clypeata 0 0 0 0 2 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 1 1 0 1 0 0 0 1 1 1 0 0 0 0 0 1 0 1 1 1 1 0 1 0 0 0 0 0 0 1 1 Trimiomelba dubia 0 0 0 3 2 0 0 0 0 1 1 1 2 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 Trimium brevicorne 0 0 0 3 2 0 1 1 0 0 1 1 2 0 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Quadrelba sp. 0 0 0 3 2 0 0 1 0 1 1 1 2 1 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Conoplectus acornus 0 0 0 1 2 0 0 1 0 1 1 1 1 1 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Rhexius sp. 0 0 0 1 2 0 0 1 0 0 1 1 1 1 0 1 0 0 0 0 0 0 1 1 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Sonoma baylessae 0 1 0 0 1 1 1 1 1 1 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1 Reichenbachia howardi 0 0 0 3 2 0 0 1 0 1 1 1 1 0 0 2 0 0 2 0 0 0 1 1 0 0 0 0 0 0 1 1 0 1 0 0 0 1 1 1 0 0 0 0 0 3 0 0 0 0 0 0 0 Pselaptrichus perfidus 0 0 0 3 1 0 0 1 0 1 1 1 1 1 0 0 0 1 1 1 0 0 1 0 0 0 0 0 0 1 1 1 0 3 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Lucifotychus cognatus 0 0 0 1 2 0 0 1 0 1 1 1 1 0 0 1 0 1 1 1 0 0 1 0 1 0 0 0 0 0 1 1 0 3 1 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0

2 simple sensory pegs, lateral angles rounded. Mandibles sickle-shaped with 6 teeth on incisor edge. Labial palpi small, 2-segmented. Maxillary palpi unmodified, MP1 minute and obliquely joined to MP2, MP2 pedunculate, narrower in basal 1/3, then slightly expanded and rounded distally, MP3 subtriangular, MP4 widest, unmodified, bearing apical palpal cone. Paired gular fovea present. Thorax: Prothorax moderately convex. Basal sulcus present, margin delimited by basolateral foveae. Mediobasal foveae absent. Prosternum bearing robust prosternal median carina, procoxal foveae present on either side of carina. Lateral mesoventral and mesocoxal foveae both present. Mesocoxae contiguous. Metacoxae conjunct, not separated by short intercoxal process of ventrite 1. Lateral metaventral foveae present, paired, metaventrite otherwise afoveate. Elytra narrower basally, then gradually wider posteriorly, bearing sutural striae with fovea. Lateroapical cleft present. Abdomen: First visible abdominal tergite bearing laterobasal foveae with weak, oval depression, margined anteriorly. Second tergite bearing similar, weaker depression and foveae. Other tergites unmodified. Second ventrite with basal carina extending between basolateral foveae, continuous. Third ventrite bearing small basolateral foveae. Other ventrites variously modified in some species as male secondary sexual characters. Male genitalia: Asymmetrical, somewhat laterally compressed. Dorsal diaphragm and internal sac visible. Parameres fixed, rotated to dorsal and ventral orientation. Median lobe sclerotized with varying combinations of spines and spicules.

FEMALE. Generally more heavy-bodied and dorso-ventrally flattened than males. Ocular canthus more prominent. Eyes usually reduced to absent (with the exception of E. dybasi ). Female genitalia: Largely membranous, comprising internalized ventrite 7 (visible as a transverse triangular strap in some specimens) and 2 internal sclerites that often appear sac-like.

Figures

Figs. 1–2. Eutyphlus similis, male. 1) Dorsal features: vs = vertexal sulcus; vf = vertexal foveae; mls = median longitudinal sulcus; laf = lateral antebasal foveae; bps = basal prosternal sulcus; lbps = longitudinal basal pronotal sulcus; shef = subhumeral elytral foveae; bef = basal elytral foveae; ds = discal stria; blf = basolateral foveae; bs = basal sulcus; mbd = median basal depression; I–V = visible tergite number; 2) Ventral features: gc = gular carina; gf = gular foveae; lpcf = lateral procoxal foveae; pslc = prosternal longitudinal carina; apsf = anteroprosternal foveae; mmsf = median mesosternal foveae; lmsf = lateral mesosternal foveae; lmtf = lateral metasternal foveae; lmcf = lateral mesocoxal foveae; blf = basolateral foveae; bls = basolateral sulcus; I–VII = visible abdominal ventrite number.