Stephanitis joceliae, Knudson & Rider & Knodel, 2021

Knudson, Alexander H., Rider, David A. & Knodel, Janet J., 2021, Two new species of Tingidae (Hemiptera: Heteroptera) from Panama, Zootaxa 4958 (1), pp. 334-344 : 339-340

Stephanitis joceliae , sp. nov.

( Figs. 3A–D , 4 )

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Diagnosis. Stephanitis joceliae can be separated from all other members of the genus, by the bucculae which project beyond the apex of the head; the long scape (0.37–0.40); by the ear shaped paranota which project apically beyond the head and pronotal hood; by the low, uniseriate median carina that is no more than half the height of the pronotal hood; and by the discoidal cell which is open apically.

Description. Generally flat, wings and paranota broadly expanded, veins infuscate, areolae hyaline. Head with moderately long, downcurved spines, adpressed to head; median spine extremely long, adpressed to head, extending to clypeus; frontal spines shorter, adpressed to head, nearly extending to clypeus. Antennae moderately long, when directed posteriorly extending to wing base; scape (segment I) long, tan, beset with a few scattered long hairs; pedicel (segment II) extremely short, tan; basiflagellomere (segment III) extremely long, two- and one-half times longer than scape, tan; distiflagellomere (segment IV) long, about one half times as long as preceding, fuscous basally, black on apical half, beset with long fine hairs. Bucculae contiguous apically, extended beyond apex of head, biseriate; rostrum moderately long, extending to metacoxae.

Pronotum punctate, ochraceous; tricarinate; median carina half as tall as pronotal hood, gradually sloping posteriorly, uniseriate; lateral carinae nearly as tall as median carina, uniseriate, subparallel, rounded in lateral view; pronotal hood narrow, moderately low, subtruncate anteriorly, comprised of two rows of cells along lateral margins, beset with short fine hairs; paranota slightly reflexed upwards, auriform, broadly expanded laterally and projecting anteriorly beyond apex of head, quadriseriate at widest; posterior pronotal projection areolate, extending to third abdominal tergite. Hemelytra broadly expanded; costal veins with minute hairs along lateral margins, hairs on basal fourth emerging from thickened projections; costal areas of hemelytra quadriseriate at base, five rows of areolae wide; subcostal areas biseriate on basal third, uniseriate throughout subcostal extensions; discoidal cells triseriate, open behind; sutural areas uniseriate. Metathoracic wings extremely small, not extending to middle of abdomen. Thorax dark ochraceous, venter beset with fine white wax; rostral laminae subparallel on pro- and mesothorax, slightly converging posteriorly, uniseriate. Legs slender, tan; femora long; tibiae slightly longer than femora; tarsi long, slender. Each ostiolar peritreme expanded laterally in posterior margin.

Abdomen light red, narrow; pygophore as long as two preceding abdominal segments; parameres slender, short, not reaching anterior edge of pygophore in repose.

Measurements. (n = 4; holotype in parentheses) Length: 3.20–3.57 (3.55), width at widest: 2.16–2.85 (2.85); height of hood: 0.32–0.36 (0.36); length of antennal segments one through four, respectively: 0.37–0.40 (0.40), 0.09–0.12 (0.12), 1.00–1.10 (1.10), and 0.61–0.70 (0.70).

Type material. Holotype: PANAMA: Chiriqui Pr.: Resve. La Fortuna , Est. Biologica. El. 3900’, 8˚ 43’ 18” N 82˚14’ 17”W, 7–9-I-2001, Yellow pan traps. M. Yoder (1 ♂ TAMU).

Paratypes: PANAMA: Chiriqui Pr.: Resve. La Fortuna , Est. Biologica. El. 3900’ N8˚ 43’ 18” W 82˚ 14’ 17”, 7–9-VIII-1999, ypt. 99/ 079, J. Schaffner & J. B. Woolly (1 ♂ TAMU) ;

Bocas del Toro: Trail off Oleoducto. 4km W Fortuna Hwy. 5-VIII-1999, el. 3640 ft, 08˚ 47’ 14” N 82˚ 12’ 50” W, J. B. Wolly 99/083 (1 ♀ TAMU); Colclé: 7.2 km NE. El Copé. 730m 08°37’N, 80° 35’ W, #140, 20 V –7 VI 1995, J. Ashe , R. Brooks, flt. Intercept trap (1 ♂ SEMC) .

All types will be deposited in their respective collections.

Etymology. This species is named in honor of eminent Brazilian heteropterist Jocelia Grazia in recognition of her many contributions to heteropteran systematics, especially of the Pentatomidae .

Plant associations. No plant associations are known.

Comments. This species was listed as Neobyrsa panamensis in an unpublished thesis ( Knudson 2018).

Figures

FIGURE 3. A–D. Stephanitis joceliae sp. nov. (A) Dorsal habitus. (B) Lateral habitus. (C) Ventral habitus. (D) Labels. Scale bar: A–C, 2.0 mm.

FIGURE 4. Distribution of new species of Tingidae in Panama.