Platynus macropterus (Chaudoir, 1879)

Larochelle & Larivière, Marie-Claude, 2021, Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2021 (864), pp. 1-96 : 6-7

Platynus macropterus (Chaudoir, 1879)

Fig. 37 , 80 , 159

Colpodes macropterus Chaudoir, 1879: 370 . Holotype: male (MNHN) labeled “Ex Musaeo Chaudoir (typed) / HOLO-

TYPE (red label; hand-written) / macropterus Chaud. Nouv. Zélande Guerin (hand-written) / Holotype (red label;

typed) / Holotype (typed) Colpodes macropterus Chd G.G. Perrault (hand-written) / G.G. Perrault det. 1988 (typed).” Anchomenus haasti Broun, 1882: 217 (redescribed in 1883: 217 and 1886: 747). Synonymized by Larochelle and Larivière

(2001: 138). Ctenognathus macropterus: Hutton 1904: 144 . Agonum (Platynus) haasti: Csiki 1931: 853 . Colpodes haasti: Johns 1980: 62 . Platynus macropterus: Liebherr 1998: 987 , 997.

Description. Body length 13.0– 14.2 mm. Head, pronotum, and elytra piceous black; abdomen dark brown; antennal segments 1–4 piceous black, 5–11 reddish; palpi reddish; femora and tibiae piceous black; tarsi reddish. Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric on elytra. Iridescence absent. Very shiny. Head. Mandibles moderately curved anteriorly. Labrum moderately transverse, slightly emarginate anteriorly. Thorax. Pronotum subdepressed, impunctate, deeply wrinkled on disc and across base, narrow, widest before middle; apex slightly emarginate; anterolateral angles poorly developed, rounded; anterior bead complete; sides strongly rounded anteriorly, strongly sinuate posteriorly; lateral beads complete; lateral depressions moderately wide anteriorly, much widened posteriorly; posterolateral angles subrectangular; laterobasal foveae very deep and wide, prolonged forward; posterior bead incomplete, obsolete medially; base emarginate. Legs. Metatarsomeres 5 setose ventrally (with short setae). Elytra. Subdepressed, not sloping down toward apex, oblong. Sides slightly rounded. Scutellar striole impunctate. Striae deep, impunctate. Intervals subdepressed; interval 3 with three shallow, minute setiferous punctures. Subapical sinuations feeble. Aedeagus. Lateral view ( Fig. 80 ): base strongly convex dorsally, with basal lobe narrow; middle moderately wide in apical half, moderately convex dorsally, strongly concave ventrally, with dorsal membranous area moderately wide in apical half and very long; apex strongly rounded dorsally, slightly concave ventrally, with extreme tip moderately wide and very long.

Material examined. 157 specimens ( AMNZ, BMNH, CMNH, JNNZ, LUNZ, MNHN, MONZ, NZAC).

Geographic distribution ( Fig. 159 ). North Island: AK, BP, CL, GB, HB, ND, RI, TK, TO, WN. South Island: BR, CO, DN, FD, KA, MB, MC, MK, NC, NN, OL, SD, SL, WD. Stewart Island.

Ecology. Lowland, montane. Epigean, arboreal. Mostly wet open forests (broadleaf, podocarp, beech), swamp forests, shrublands, scrublands, forest edges, and tree plantations (pine); tussock grasslands and pastures. Associated with streams. Mostly open ground. Nocturnal; hides during the day under logs and stones.

Biology. Seasonality: September–March, May, July–August. Tenerals: November–January, March. Predacious (based on mouthpart morphology). Predators: Feral cats. Occasionally infested with mites and fungi (Laboulbeniales). Defense mechanism: when alarmed, the adult emits a burnt chocolate-like smell.

Dispersal power. Macropterous. Occasional flier to artificial lights. Swift runner. Occasional climber (on trees). Favored by human activities.

Collecting techniques. Turning logs and stones.

References. Larochelle and Larivière 2001: 138–139 (catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 114 (list), 2016: 38 (list).

Figures

Figures 37–40. Dorsal habitus. 37) Platynus macropterus. 38) Notagonum chathamense. 39) N. lawsoni. 40) N. feredayi. Scale line = 1 mm.

Figures 80–86. Aedeagus, lateral. 80) Platynus macropterus. 81) Notagonum chathamense. 82) N. lawsoni. 83) N. feredayi. 84) N. submetallicum. 85) Prosphodrus mangamuka new species. 86) P. waltoni.

Figures 154–159. Species distribution maps. 154) Maoriplatynus marrisi new species. 155) Notagonum chathamense. 156) N. feredayi. 157) N. lawsoni. 158) N. submetallicum. 159) Platynus macropterus.