Titiotus shasta, Platnick & Ubick, 2008

Platnick, Norman I. & Ubick, Darrell, 2008, A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae), American Museum Novitates 3608 (1), pp. 1-36: 10-14

publication ID

http://doi.org/ 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2

persistent identifier

http://treatment.plazi.org/id/038F87AE-FFB7-FFF5-FF7D-F9F0A5E25A84

treatment provided by

Carolina

scientific name

Titiotus shasta
status

new species

Titiotus shasta   , new species

Figures 17–21 View Figs ; map 1

TYPE: Male holotype from McArthur , Shasta Co.   , California (June 18, 1942; W. Pearce), deposited in   AMNH.

ETYMOLOGY: The specific name is a noun in apposition to the genus name taken from the type locality.

DIAGNOSIS: Males resemble those of T. californicus   and T. hansii   but have a much smaller ventral branch on the middle prong of the tibial apophysis (fig. 19); females resemble those of T. californicus   but have a posteriorly narrowed epigynal septum (fig. 20).

MALE: Total length 10. Coloration as in T. californicus   . Leg spination: femora: I, II r1-2- 1; III, IV p1-2-1; tibiae: I, II d1-0-1, v4-4-4, r1- 1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2; IV r1-1-2. Embolus expanded prolaterally, beak-shaped distally; median apophysis with elongate tip, with only tiny subdistal projection; middle prong of tibial apophysis bifid, with ventral branch much smaller than dorsal branch (figs. 17–19).

FEMALE: Total length 12. Coloration as in T. californicus   . Leg spination: femora: I, II r1- 2-1; III r2-1-1; IV p2-0-1; tibiae: I v4-4-4, r0-1- 0; II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I p0-1-0, r0-1-0; II p1-1-0, r0-1-0; III p1- 2-2. Epigynal septum wide, occupying most of epigynal width, except posteriorly, where narrowed; posterior ducts elbowed (figs. 20, 21).

OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Colusa Co.: Deafy ( Dixie ) Glade Campground on Forest Road , 13 mi W Stonyford, Mendocino National Forest, 39 ° 21 9 N, 122 ° 42 9 W, no date, elev. 1000 m (J. Schweikert, CAS) GoogleMaps   , 13. Glenn Co.: Forest Road, 5.5 mi S County Road 308, 9 mi SW Elk Creek, Mendocino National Forest , 39 ° 30 9 N, 122 ° 39 9 W, Mar. 30, 1997, elev. 1100 m (J. Schweikert, CAS) GoogleMaps   , 13. Lake Co.: Lucerne, Aug. 17, 1958, molasses bait trap (H. Leech, CAS), 1♀. Mendocino Co. : 7.5 km NE Covelo, 28201 Mendocino Pass Road , 39 ° 50 9 N, 123 ° 10.7 9 W, Sept. 2, 1996, elev. 1680 ft (C. Prince, CAS), 1♀. Shasta Co.   : Lake Shasta , June 14, 2006 (C. Abreu, UCR)   , 13; Palo Cedro , June 17, 2006 (D. McConnel, UCR)   , 23; Redding , Aug. 26, 2004 (A. Tennett, UCR), 1♀, June 2, 2006, in bathtub (A. Olivas, UCR), 1♀, June 4, 2006 (H. Ray, UCR)   , 13, June 16, 2006 (G. Snook, UCR)   , 13, same date (W. Mapes, UCR), 1♀, June 28, 2006 ( T. Ressler, UCR), 1♀, June 29, 2006 (M. Wooden, UCR), 1♀. Trinity Co.: Hayfork Ranger Station , May 19, 1973 (M. Bentzien, UCB)   , 13.

DISTRIBUTION: Known only from northwestern California (map 1).

Titiotus flavescens (Chamberlin and Ivie)  

Figures 22–26 View Figs ; map 2

Liocranoides flavescens Chamberlin and Ivie, 1941: 24   , fig. 18 (female holotype from Ben Lomond , Santa Cruz Co., in AMNH, examined).

Titiotus flavescens: Lehtinen, 1967: 271   .

DIAGNOSIS: Males resemble those of T. shantzi   in having the middle prong of the tibial apophysis relatively narrow and the dorsal prong relatively short, but differ in having the middle prong more deeply incised (fig. 24); females have a distinctive epigynal septum, which is greatly narrowed posteriorly (fig. 25).

Map 2. Central California, showing distributions of eight species of Titiotus   .

MALE: Total length 11. Coloration as in T. californicus   . Leg spination: femora: I, II r1-2- 1; IV p0-2-1, r0-1-1; tibiae: I, II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I p1-0-0, r0-1-0; II p1-1-0, r1-1-0; III p1-2-2. Embolus with subapical, distally directed projection; median apophysis with subdistal projection; middle prong of tibial apophysis small, bifid, dorsal branch distally invaginated, dorsal prong relatively short (figs. 22–24).

FEMALE: Total length 14. Coloration as in T. californicus   , except proximal leg segments with dorsal dusky markings. Leg spination: femora: I, II r1-2-1; III, IV p1-2-1; tibiae: I p1- 0-0, v4-4-4, r1-0-0; II v4-4-4, r0-2-0; III, IV p1- 0-1, r1-0-1; metatarsi: III p1-2-2. Posterior quarter of epigynal septum greatly narrowed, forming stalk; posterior ducts oblique (figs. 25, 26).

MATERIAL EXAMINED: United States: CALI- FORNIA: Santa Clara Co.: 27950 N Carnegie Ave., Santa Clara, June 8, 1993 ( T. Carb, AMNH), 13; San Jose , Oct. 21, 1974, in house (D. Ubick, CDU)   , 13. Santa Cruz Co.: Aptos , July 10, 1946 (E. Ross, AMNH)   , 1♀; Bat Cave, Santa Cruz , Apr. 21, 1979 (D. Rudolph, Cowan, Van Ingen, AMNH)   , 23; Ben Lomond , May 1934 (L. Saylor, AMNH)   , 1♀ (holotype), June 25, 1952 (W. Gertsch, AMNH, JCC)   , 13, 21♀, July 4, 1958 (W. Gertsch, V. Roth, AMNH), 12♀, Apr. 3, 1960 (W. Gertsch, W. Ivie, R. Schrammel , AMNH), 93, 11♀, Sept. 23, 1961 (W. Gertsch, W. Ivie, AMNH)   , 53, 9♀; Boulder Creek , Aug. 1, 2004, in house ( T. Nygard, UCR), 1♀, Oct. 7, 2004, in house (J. Sanchez, UCR)   , 13; 1 mi NE Boulder Creek , June 29, 1989, dry redwood forest, elev. ca 800 ft (A. Sundman, CDU)   , 1♀; Cave Gulch , Nov. 16, 1979, redwood forest (D. Ubick, CDU)   , 1♀; Cave Gulch, cave near Univ. of California, Santa Cruz , Nov. 4, 1972 ( T. Stearns, CAS), 1♀; Cave Gulch , IXL Cave, Dec. 6, 1980, on cave wall ( T. Briggs, CAS), 13; Felton, Aug. 16, 1959 (W. Gertsch, V. Roth, AMNH), 2♀, Apr. 3, 1960 (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 1♀, June 1994 (S. Tjosvold, CAS)   , 1♀, Sept. 2001 ( R. Knight, UCR), 13, Oct. 3, 2004, in shoe in bedroom (M. Mister, UCR)   , 13; N Soquel, Oct. 6, 2004 (A. Fournier, UCR)   , 13.

DISTRIBUTION: Known only from Santa Clara and Santa Cruz counties south of the San Francisco Bay (map 2).

AMNH

American Museum of Natural History

CAS

California Academy of Sciences

UCR

University of California

T

Tavera, Department of Geology and Geophysics

UCB

University of California at Berkeley

V

Royal British Columbia Museum - Herbarium

R

Departamento de Geologia, Universidad de Chile

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Zoropsidae

Genus

Titiotus

Loc

Titiotus shasta

Platnick, Norman I. & Ubick, Darrell 2008
2008
Loc

Titiotus flavescens:

Lehtinen, P. T. 1967: 271
1967
Loc

Liocranoides flavescens

Chamberlin, R. V. & W. Ivie 1941: 24
1941