Titiotus heberti, Platnick & Ubick, 2008

Platnick, Norman I. & Ubick, Darrell, 2008, A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae), American Museum Novitates 3608 (1), pp. 1-36: 20-24

publication ID

http://doi.org/ 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2

persistent identifier

http://treatment.plazi.org/id/038F87AE-FFAD-FFEF-FD42-F996A4C15D96

treatment provided by

Carolina

scientific name

Titiotus heberti
status

new species

Titiotus heberti   , new species

Figures 47–51 View Figs ; map 1

TYPES: Male holotype and female allotype from Wilsonia, Kings Canyon National Park , Tulare Co.   , California (Sept. 13, 1959; W. Gertsch, V. Roth)   , deposited in AMNH.

ETYMOLOGY: The specific name is a patronym in honor of Mr. Blaine Hebert, assiduous collector of these and other Californian spiders.

DIAGNOSIS: Males resemble those of T. tulare   in having a large, wide dorsal prong on the tibial apophysis, but differ in having a distally narrower middle prong (fig. 49); females differ from those of T. tulare   in having a posteriorly wider epigynal septum (fig. 50).

MALE: Total length 12. Coloration as in T. californicus   . Leg spination: femora: I p0-3-1, r1-2-1; II r1-2-1; IV p1-2-1; tibiae: I, II d1-0-1, r1-1-0; III r1-0-1; metatarsi: I p1-0-0, r1-1-0; III p1-2-2. Embolus twisted near base, distally wide; median apophysis with subdistal and retrolateral projections; middle prong of tibial apophysis bifid, with subequal branches, dorsal prong large, wide, flattened (figs. 47– 49).

FEMALE: Total length 17. Coloration as in T. californicus   . Leg spination: femora: I p0-3- 1, r1-2-1; II r1-2-1; III r1-1-2; IV p1-2-1; patella III p0-0-0; tibiae I, II r1-1-0; metatarsi III, IV p1-2-2. Epigynal septum wide, with posteromedian bulge occupying most of septal width; posterior ducts oblique, arched (figs. 50, 51).

OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Fresno Co.   : Cedar Grove, Kings River Canyon , July 16, 1952, elev. 4633 ft (W. Gertsch, AMNH), 8♀   , July 5, 1956, elev. 4633 ft (W., J. Gertsch, AMNH), 23, 6♀   ; Hume Lake , Dec. 17, 1984 (K. Walker, CAS), 1♀   ; Lily Pond , 2 mi W Wishon Lake, Oct. 31, 1982, in sandy soil (M. Marshall, CAS), 1♀   ; Ranger Station, Kings River, Kings Canyon National Park , June 17, 1969, in house (M. Zardus, CAS), 13; Shaver Lake , July 15, 1952 (W. Gertsch, AMNH), 5♀   , Sept. 12, 1959 (W. Gertsch, V. Roth, AMNH), 123, 3♀   ; vicinity Wishon Resv. , July 2–6, 1969 (D. Marqua, MET), 13, 2♀   , May 28–June 14, 1972 (M. Thompson, MET), 2♀   . Kern Co.: Cedar Creek Camp Ground, 0.25 mi N Highway 155, Greenhorn Mountains , 35 ° 44.44 9 N, 118 ° 35.54 9 W, Aug. 25, 2006, on wall of old mine shaft in steep, rocky canyon side near stream, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀   , Aug. 26, 2006 (matured Sept. 4, 2006), on ceiling at mouth of old mine shaft in steep, rocky canyon side stream nearby conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀   , Aug. 26, 2006 (matured Sept. 8, 2006), on base of tree near stream in steep-sided canyon, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀   , Aug. 26, 2006 (matured Dec. 9, 2006), on side of big rock near stream in steep-sided canyon, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 13. Madera Co.   : Cascadel Woods, T   8S,R 23E,S16, July 1–3, 1983 (J. Aotok, CAS), 13; Coarsegold, no date (Oliva, UCR), 1♀   , Apr. 21, 2004 (B. Harrigom, UCR), 1♀   ; Wishon, July 5, 1971 (D. Marqua, MET), 13. Mariposa Co.: Foresta Road , Yosemite National Park , 37 ° 42.242 9 N, 119 ° 43.989 9 W, June 15–18, 2003, pitfalls, elev. 1392 m (A. Leache, C. Conroy, UCB), 63. Tulare Co.   : Ash Mountain , Sequoia National Park , Apr. 26, 1951 (E. Schlinger, AMNH), 13; near Ash Mountain , Sequoia National Park , July 9, 1958 (W. Gertsch, V. Roth, AMNH), 13, 5♀   ; 1 mi below Belnap Springs, near Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 23, 2♀   ; Carmoe Crevice , Nov. 3, 1993 (J. Krejca, CAS), 13; NE Case Mountain Peak   , 36.41790 ° N, 118.7933 ° E, July 14–18, 2003, dry pitfall, elev. 1691 m ( R. Fisher, AMNH), 13; NE Case Mountain Peak GoogleMaps   , 36.41870 ° N, 118.79264 ° E, July 14–18, 2003, dry pitfall, elev. 1672 m ( R. Fisher, AMNH), 13; NNE Case Mountain Peak, E of E end of Salt Creek GoogleMaps   , 36.42059 ° N, 118.79776 ° E, May 2003, dry pitfall, elev. 1749 m ( R. Fisher, AMNH), 13 GoogleMaps   , 36.42083 ° N, 118.79163 ° W, Sept. 2002, dry pitfall, elev. 1609 m ( R. Fisher, AMNH), 1♀ GoogleMaps   ; NW Case Mountain Peak, S of Salt Creek   , 36.41871 ° N, 118.81930 ° W, May 2003, dry pitfall, elev. 1709 m ( R. Fisher, AMNH), 13; just S Case Mountain Peak GoogleMaps   , 36.407 ° N, 118.8016 ° E, July 14–18, 2003, dry pitfall, elev. 1998 m ( R. Fisher, AMNH), 13; SE Case Mountain Peak, Salt Creek Ridge GoogleMaps   , 36.39964 ° N, 118.77975 ° W, Sept. 2002, dry pitfall, elev. 1808 m ( R. Fisher, AMNH), 13; W Case Mountain Peak GoogleMaps   , 36.41120 ° N, 118.80909 ° W, July 2002, dry pitfall, elev. 1951 m ( R. Fisher, AMNH), 1♀ GoogleMaps   , 36.41139 ° N, 118.80843 ° W, July 2002, dry pitfall, elev. 1932 m ( R. Fisher, AMNH), 1♀ GoogleMaps   , July 14–18, 2003, dry pitfall, elev. 1951 m ( R. Fisher, AMNH), 1♀   ; just W Case Mountain Peak   , 36.41139 ° N, 118.80843 ° W, July 14–18, 2003, dry pitfall, elev. 1932 m ( R. Fisher, AMNH), 13; Cedar Cave, Redwood Canyon, Kings Canyon National Park , Aug. 16, 1984, under rock outside cave entrance, elev. 1,600 m (D. Ubick, CDU), 1♀ GoogleMaps   ; Crystal Sequoia Cave, Sequoia National Park , Apr. 16, 1946 (F. Oberhansley, AMNH), 1♀   , July 26, 1991 (D. Ubick, CDU), 1♀   ; cave in bank on road to Crystal Cave, Sequoia National Park , Nov. 6, 1949, in total darkness (A. Lange, AMNH), 1♀   ; Harry’s Bend Cave, July 10, 2004 (J. Krejca, A. Gluesenkamp, C. Richard, CAS), 13; Kaweah Power Station #3, Ash Mountain   , 40 mi NE Visalia, July 6, 1983 (D. Burdick, CAS, CDU), 23, May 9, 1993 (D. Burdick, CAS), 13, July 2, 1995 (D. Burdick, CAS), 13; Lindcove , July 13, 1976, dead on doorstep (D. Carroll, UCR), 1♀   ; McIntire Creek, near Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 13, 1♀   ; 5–10 mi on Mineking Road from Three Rivers , July 4, 1956, elev. 2000–4000 ft ( AMNH), 13, 4♀   ; Moorhouse Creek, near Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 13   ; 1 mi W Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 2♀   ; Roads End, Kern River, July 3, 1956 (W. Gertsch, AMNH), 13; Soda Creek , W Camp Nelson , July 11, 1958 (W. Gertsch, V. Roth, AMNH), 23, 2♀   ; Wilsonia, Kings Canyon National Park , Sept. 13, 1959 (W. Gertsch, V. Roth, AMNH), 23, 3♀   ; 9 mi N Woodlake, Mar. 22, 1941 (S., D. Mulaik, AMNH), 13. Tuolumne Co.: Tuolumne , May 2004, in house (M. Rauscher, CDU), 13   .

DISTRIBUTION: Known only from the Sierras, generally to the south of T. gertschi   (map 1).

V

Royal British Columbia Museum - Herbarium

AMNH

American Museum of Natural History

CAS

California Academy of Sciences

T

Tavera, Department of Geology and Geophysics

UCR

University of California

UCB

University of California at Berkeley

R

Departamento de Geologia, Universidad de Chile

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Zoropsidae

Genus

Titiotus