Titiotus humboldt, Platnick & Ubick, 2008

Platnick, Norman I. & Ubick, Darrell, 2008, A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae), American Museum Novitates 3608 (1), pp. 1-36 : 28-30

Titiotus humboldt , new species Figures 72–76 ; map 3

TYPES: Male holotype and female allotype taken 10 mi east of Bridgeville , Humboldt Co. , California (Aug. 20, 1959; W. Gertsch), deposited in AMNH.

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: This species resembles T. marin and T. costa in having the ventral prong of the tibial apophysis relatively short and wide, and situated close to the base of the middle prong; males can be distinguished by having the third most ventral prong of the tibial apophysis relatively short, only slightly expanded distally, and not incised (fig. 74), females by the relatively wide epigynal septum and protuberant posterolateral epigynal margins (fig. 75).

MALE: Total length 11. Coloration as in T. californicus . Leg spination: femora: I p1-2-1; II r1-2-1; IV p1-2-1; tibiae: I d1-0-1, v4-4-4, r0-1- 0; II d1-0-1, p1-0-0, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Embolus prolaterally expanded at base, distally narrow; median apophysis with only tiny subdistal projection; two most ventral prongs of tibial apophysis small, third relatively short, square (figs. 72–74).

FEMALE: Total length 12. Coloration as in T. californicus . Leg spination: femora: I, II r1- 2-1; III p2-1-2, r1-1-3; IV p2-1-1; tibiae: I v4-4-

74. Same, retrolateral view. 75. Epigynum, ventral view. 76. Same, dorsal view.

4, r1-0-0; II v4-4-4, r1-1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Epigynal septum wide, wider anteriorly than posteriorly, posterolateral epigynal margins protuberant; posterior ducts widely separated (figs. 75, 76).

OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Humboldt Co. : Five Mile Creek at junction of Highway 96, 4.2 mi S Somes Bar, Oct. 2, 1979, on wall of 6 ft diameter conduit under highway, elev. 450 ft Saussure , AMNH), 1♀, Apr. 9, 1979 (D. Rudolph, B. Martin, S. Winterath, AMNH) , 13; Samwell Cave, 40 ° 55 9 N, 122 ° 14 9 2 0 W, June 7, 1995, elev. 1450 ft (D. Ubick, CDU), 1♀, Aug. 30, 1996, elev. 1450 ft (D. Ubick, CDU), 1♀; Shasta Lake Caverns, N side Shasta Lake , 16 mi NNE Redding, Apr. 10, 1979 (D. Rudolph, B. Martin, S. Winterath, AMNH), 1♀. Siskiyou Co.: Ney Springs , 5 mi W Mount Shasta, Sept. 2, 1959 (W. Gertsch, V. Roth, AMNH), 1♀. Tehama Co. : Woodson Bridge State Recreation Area , 7 mi E Corning, Aug. 29, 1996, oak camp, at night (D. Ubick, CDU), 1♀. Trinity Co. : Big Canyon , 2.6 mi E Camp Trinity, Aug. 12, 1990, under rock (D. Ubick, CDU), 1♀; Del Loma, N fork, Trinity River , July 25, 1953 (W., J. Gertsch, AMNH), 15♀; Forest Glen Cave , on open wall, dry, 75 ft inside cave ( R. Graham, AMNH), 23; Weaverville to 10 mi E Weaverville, Apr. 6, 1960 (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 23 .

DISTRIBUTION: Northwestern California (map 3).

Map 3. Northern California, showing distributions of three species of Titiotus .

(W. Icenogle, WRI), 1♀. Mendocino Co.: near Headquarters , Hopland Field Station , 4 mi E Hopland, Apr. 6, 1972, matured Apr. 28, 1972 (M. Bentzien, UCB), 13. Shasta Co. : Castle Crags State Park , 4 mi W Castella, July 26, 1953 (W., J. Gertsch, AMNH), 1♀ ; Hazel Creek , July 4, 1952 (W. Gertsch, AMNH), 23, 4♀ ; Potter Creek Cave , 40 ° 47 9 00 0 N, 122 ° 16 9 45 0 W, Nov. 24, 1950 (N. Bostick, AMNH), 13, Sept. 7, 1961 (W. Gertsch, AMNH), 1♀ ; Samwell Cave , 25 mi N Redding, no date ( R. de Saussure, AMNH), 1♀ , Apr. 28, 1959, at entrance ( AMNH), 1♀ , June 5, 1959 ( R. de

Figures

Figs. 72–76. Titiotus humboldt, new species. 72. Left male palp, prolateral view. 73. Same, ventral view.