Timbellus longicanalis, Merle & Pacaud & Ledon & Goret, 2024

Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard, 2024, New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe, Geodiversitas 46 (15), pp. 495-551 : 514-516

Timbellus longicanalis n. sp.

( Figs 11 A-D; 26E, F)

urn:lsid:zoobank.org:act:59D8D20D-41AD-43F9-8A3B-B247A150B335

Timbellus fusoides – Merle et al. 2011: 446, pl. 101 fig. 5a, b only [non Deshayes, 1865].

TYPE MATERIAL. — Holotype. France • Normandy, Manche, Fresville ; Calcaire de Fresville ; middle Eocene (uppermost Lutetian); MNHN.F.A24101 (Le Marchand coll.), H: 35 mm ( Figs 11A, B ; 26E ). Paratype. France • 1 spm; same as for the holotype; MNHN.F.A24100 (Le Marchand coll.) ( Figs 11C, D ; 26F ).

ETYMOLOGY. — From the Latin adjective longus meaning elongated and canalis meaning canal, refecting the elongated siphonal canal of this species.

TYPE HORIZON. — Calcaire de Fresville, middle Eocene (uppermost Lutetian), see Bignot et al. (1968).

TYPE LOCALITY. — France, Normandy, Manche, Fresville, see APGN (2024).

DISTRIBUTION. — Normandy ( Calcaire de Fresville ); middle Eocene (uppermost Lutetian).

DESCRIPTION Protoconch not preserved. Teleoconch up to 35 mm in height, up to 14 mm in width, fusiform in profile, composed of six whorls. Moderately high spire. Last whorl up to 77% of total length. Apical angle 41° excluding spine and 61° including spines. Spiral sculpture with moderately marked primary cords on varices. First and second whorl eroded. Third whorl: appearance of weak s1 and P2 on intervarices. Fourth whorl: appearance of weak P1 cord. Fifth whorl: no change. Sixth whorl: P1-P4 marked, P5-P6 weak, s1, s2, s6, P6 not atrophied, s1-s2, s6 weak; ADP, MP and ABP weak. Axial sculpture eroded on two early whorls. From third last whorl: three varices and one intervarix intercalated. Higher relief of intervarices on P2. Varices lamellose, slightly winged. Appearance of P1 spine on third whorl, no other spines adapically. On last whorl P1 spine short, straight, slightly bent adaxially. On last whorl growth lamellae erect giving scabrous aspect to shell surface. Ovate aperture up to 29% of diameter and up to 78% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly, forming very slightly curved inductura at base. Parietal lip adherent. Outer lip with simple, weak denticles including ID, D1 to D6. Siphonal canal narrow, open, straight, up to 59% of apertural length. Pseudoumbilicus narrow.

COMPARISONS

This species was previously identified as Timbellus fusoides ( Deshayes, 1865) by Merle et al. (2011: pl. 101, fig. 5a, b). Nevertheless, attribution to Murex fusoides is problematic, as the type material on which that Bartonian species is based is poorly preserved. It corresponds to two syntypes housed in the collection of the University Lyon (no. UCBL-EM 33383-84; Fig. 11E ). The largest is a strongly eroded specimen with most of its axial sculpture, particularly its winged varices lost. This loss of sculpture gives the shell an artificially fusiform shape. The second specimen is smaller and also strongly eroded, but P1 spine is rather turned adapically as in some Timbellus tripteroides ( Lamarck, 1822) and its aperture resembles that of T. tripteroides . We designate herein this smaller specimen (UCBL-EM 33384; Fig. 11F, G ) as the lectotype of M. fusoides . Therefore, M. fusoides can be regarded as a possible junior synonym of T. tripteroides or a nomen dubium as it is impossible to clearly identify a species based on the type material. Moreover, Bartonian specimens from the Paris Basin identified as T. fusoides are present in several collections housed in the MNHN (e.g., Faullummel, Pons and Schtrock colls). Examination of these specimens shows them to represent eroded T. tripteroides (probably reworked from the Lutetian) or to large and eroded Pterochelus contabulatus ( Lamarck, 1803) . The type material of T. longicanalis n. sp. comes exclusively from the upper Lutetian of Normandy. It is very well preserved and its delicate and winged varices are not eroded. Thus, its fusoid shape is not due to a post-mortem alteration. The species differs from T. tripteroides by having more staged and more quadrate whorls, more adaxially oriented P1 spines, a straighter siphonal canal, a less rounded aperture, and by having weaker internal denticles and spiral cords.

Figures

FIG. 11. — Spiral cords and internal denticles of the outer lip in Timbellus longicanalis n. sp. (A-D) and type material of Murex fusoides (Deshayes, 1865) (E-I): A, B, holotype,MNHN.F.A24101 (Le Marchand coll.),Fresville, Manche,Normandy, France,middle Eocene (upper Lutetian);C, D, paratype,MNHN.F.A24100, Fresville;E, type material of M. fusoides in the Deshayes’s collection corresponding to two eroded T. tripteroides; F, G, lectotype designated herein,UCBL-EM 33384, Valmondois, Val d’Oise, Paris Basin, France, middle Eocene (Bartonian); H, I, paralectotype, UCBL-EM 33383, Sainte-Aulde (Caumont), Seine-et-Marne, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A-D, P. Loubry (MNHN/CNRS); E-I, E. Robert (UCBL).

FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm.