Scolytocis howdeni Lopes-Andrade

Lopes-Andrade, Cristiano, 2008, An essay on the tribe Xylographellini (Coleoptera: Tenebrionoidea: Ciidae), Zootaxa 1832 (1), pp. 1-110: 21-22

publication ID 10.11646/zootaxa.1832.1.1

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scientific name

Scolytocis howdeni Lopes-Andrade

sp. nov.

Scolytocis howdeni Lopes-Andrade   sp. nov. ( Figs 39–41 View FIGURES 39–41 , 58 View FIGURES 57–59 , 64B View FIGURES 63–65 , 67B View FIGURES 66–68 , 72 View FIGURES 72–73 , 80 View FIGURE 80 )

Etymology The species is named after Henry F. Howden, who collected six specimens of the type series.


The species belongs to the danielssoni   species-group, and so it is distinguishable from the other Scolytocis species   by the biconcave prosternum with a narrow longitudinal carina at midline, the rugose border along the posterior pronotal margin and the comparatively large tibiae. It differs from the other species in the group mainly by the close pronotal punctation, with punctures separated by a distance of one puncture width or less.


Holotype. ( Figs 39–41 View FIGURES 39–41 ) Measurements in mm: TL 2.00; PL 0.79; PW 0.89; EL 1.21; EW 0.95; GD 0.89. Ratios: PL/PW 0.88; EL/EW 1.28; EL/PL 1.53; GD/EW 0.94; TL/EW 2.11. Body dark brown, head and pronotum slightly darker than elytra; basal antennomeres, mouthparts and tarsi yellowish brown; dorsal and ventral surfaces almost entirely glabrous. Head with dorsum coarsely punctate; punctures irregular, with interstice finely microreticulate. Each antenna ( Fig. 64B View FIGURES 63–65 , paratype) with nine antennomeres (FL 0.098mm; CL 0.160mm; CL/FL 1.64); length of the antennomeres (in mm) as follows: 0.118; 0.055; 0.053; 0.015; 0.015; 0.015; 0.050; 0.045; 0.065. Eyes with greatest width 1.33X the basal width of scutellum. Pronotum coarsely punctate; punctures irregular, limits distinct, usually separate by a distance of one puncture width or less, but not coalescent; interstice between punctures finely microreticulate; anterolateral margins straight; posterolateral angles broadly rounded; posterior margin with a conspicuous rugose border along it. Scutellum rugose, basal width 0.13X the EW. Elytra with punctation slightly coarser, denser and more irregular than that of pronotum; interstice between punctures smooth; elytral apex truncate, with a small concavity when seen from above; cuticular globules absent. Each hindwing ( Fig. 58 View FIGURES 57–59 ) with apical area bearing one vague, incomplete pigmented line near the anterior margin. Prosternum biconcave, with a narrow longitudinal carina at midline; surface microreticulate to striated. Metaventrite glabrous, surface microreticulate to striated; discrimen indistinct; either side bearing large obsolescent punctures. Abdominal ventrites glabrous; surface microreticulate. Each metatibia ( Fig. 67B View FIGURES 66–68 ) near 3.0X as long as broad; outer apical and outer lateral edges forming an obtuse angle that is broadly rounded; outer edge with spines (around 20) regularly distributed and more close to each other at apical third (outer apical edge), then getting sparser. Male genitalia (in a paratype). ( Fig. 72 View FIGURES 72–73 ) Apex of tegmen (sclerotized apical half) subtriangular, base membranous; median lobe membranous, elongate, near 1.2X longer than tegmen.

Type series

Holotype. (CNCI) Mexico: / 3 mi. N.W. San Cristobal L.C., Chis. Mex. V. 29 1969 H.F. Howden / Scolytocis howdeni Lopes-Andrade   HOLOTYPUS [printed on red paper]/. Paratypes. Mexico: 4 specimens (2 CNCI, 2 LAPC), same data as holotype; 1 specimen (ANIC) / 6.4mi. S. Tenango de Doria Hidalgo MEX. July 24, 1969 / S. & J. Peck Collectors / Ganoderma sp.   / J. F. Lawrence Lot [printed] 3079 [handwritten]/. All the paratypes with an additional label / Scolytocis howdeni Lopes-Andrade   PARATYPUS [printed on yellow paper]/


Measurements in mm (n = 6, including the holotype): TL 1.84–2.00 (1.92 ± 0.07); PL 0.68–0.79 (0.70 ± 0.04); PW 0.79–0.89 (0.85 ± 0.04); EL 1.16–1.26 (1.20 ± 0.04); EW 0.84–0.95 (0.92 ± 0.04); GD 0.84–0.89 (0.86 ± 0.03). Ratios: PL/PW 0.76–0.88 (0.83 ± 0.04); EL/EW 1.22–1.38 (1.31 ± 0.06); EL/PL 1.53–1.85 (1.72 ± 0.11); GD/EW 0.89–1.00 (0.93 ± 0.04); TL/EW 1.94–2.19 (2.09 ± 0.08).

Distribution Known from Chiapas and Hidalgo, Mexico ( Fig. 80 View FIGURE 80 ).

Host fungi Ganoderma sp.   ( Ganodermataceae   ).


It resembles Scol. indecisus   sp. nov. in the coarse and dense punctation of elytra and pronotum, but is easily distinguished by the features mentioned in the diagnosis.