Agyneta tuberculata, Dupérré, Nadine, 2013

Dupérré, Nadine, 2013, Taxonomic revision of the spider genera Agyneta and Tennesseellum (Araneae, Linyphiidae) of North America north of Mexico with a study of the embolic division within Micronetinae sensu Saaristo & Tanasevitch 1996, Zootaxa 3674 (1), pp. 1-189 : 115

Agyneta tuberculata new species

Figs 368–375 , map 21

Type material: Male holotype from Arizona, Cochise county, Portal, Sept. 10, 1981, in house, W.J. Gertsch collector ( AMNH). EXAMINED.

Etymology: The specific name is an adjective in apposition taken from the Latin in reference to the heavily tuberculate cymbium.

Diagnosis: Males are diagnosed from all Agyneta species by their tricorn dorsal cymbial tubercle and their large ventral tubercle with protuberance ( Fig. 370 , arrow). Females are diagnosed from all Agyneta by their narrow, triangular epigynal slits ( Fig. 373 ). For differentiation from A. crista see diagnosis of the latter.

Description: Male: Total length 2.02; carapace length 0.87, width 0.72.

CEPHALOTHORAX: Carapace orange, shiny, finely reticulate; lightly suffused with gray along margin, radiating lines; trident mark sometimes present. Sternum suffused with dark gray. Clypeus height 2. Chelicerae orange, apical part lighter, excavated; ~15 seta-tipped tubercles; promargin three denticles, retromargin two tiny denticles. Cheliceral stridulatory organ visible ~58 striae well spaced, getting closer basally. ABDOMEN: Uniformly light to dark gray. LEGS: Orange, distal half of femur slightly darker; leg I total length: 4.11; leg III total length: 2.74; Tm I: 0.15, Tm IV: absent. GENITALIA: Palpal retrolateral tibial apophysis long, tip slightly rugose; dorsal tibial apophysis wide, smooth; two retrolateral trichobothria and one dorsal ( Fig. 368 ). Cymbium triangular; glabrous depression present ( Fig. 368 ); dorsal cymbial tubercle with two rounded tip, carinated and smooth; ventral tubercle long, smooth and rounded, with extra process basally ( Figs 369, 370 ); prolateral notch medium ( Fig. 369 ). Paracymbium apical pocket medium, anterior pocket long making a large cover, posterior pocket absent ( Fig. 368 ). Embolus tip wide and pointed, with wide and dentate, prong; ventrally rugose; Fickert’s gland absent; ventral lamella transparent, spiny; thumb reaching over embolus proper ( Fig. 371 ). Embolus proper set apically, dorsal part slightly larger ( Fig. 371 ). Anterior terminal apophysis narrow with short protrusions; posterior terminal apophysis narrow, with one small point midway and one small spur basally ( Fig. 372 arrow); lamella characteristica wide, well sclerotized with numerous short spines, one large prong medially and rounded tip ( Fig. 372 ).

Female: Total length 1.97; carapace length 0.92, width 0.60.

CEPHALOTHORAX: Coloration as male. Chelicerae orange; promargin five teeth, retromargin three denticles. Cheliceral stridulatory organ visible ~50 striae, narrowly spaced throughout. ABDOMEN: Same as male. LEGS: Same as male; palpal segment slightly suffused with gray, tarsal claw absent; leg I total length: 3.13; leg III total length: 2.20; Tm I: 0.20, Tm IV absent. GENITALIA: Epigynum with wide proximal part of scape, almost rectangular; epigynal slits narrow and triangular; pit hook depression deep ( Fig. 373 ); lateral lobes elongated; stretcher and pit medium ( Fig. 374 ). Median part of scape long and narrower apically; genital pores situated at base of lateral lobes pockets ( Fig. 375 ). Internal genitalia with large and sinuous receptacula ( Figs 374, 375 ).

Other material examined: USA: Arizona: Catalina Mountains, Molino Basin, 27.xi.1976, 13, V. Roth, Schroepfer ( AMNH); Portal, 10.x.1981, in house, 13, W. Gertsch ( AMNH). Nebraska: Hasley, 12–19. viii.1957, 13, H. Hemzlik ( AMNH). New Mexico: T65 R10E S36, 01.vii.1972, 13, Riechert ( AMNH); West of Silver City, 08.ix.1941, 1Ƥ, W. Ivie ( AMNH). Texas: Edinburg, 10.i.1939, 13, S. Mulaik ( AMNH); Mission, 1Ƥ, S. Mulaik ( AMNH); New Deal, 21.vii.1972, 13, J. Rowland ( AMNH); Raven Ranch, vi.1941, 1 Ƥ, S. Mulaik ( AMNH); Rio Grande City, 29.iv.1939, 13, S. Mulaik ( AMNH); no specific locality, i–iii.1936, 2 Ƥ, L. Davis ( AMNH). Utah: Bluff, 10.v.1933, 132Ƥ ( AMNH); Little Castle Valley, 30.x.1931, 43, W. Ivie ( AMNH).

Distribution: Southwest USA, north to Nebraska.

Figures

FIGURES 368 – 375. Agyneta tuberculata n. sp. 368 – 372, Male. 373 – 375, Female. 368. Palp, retrolateral view. 369. Palp, prolateral view. 370. Variation of dorsal and ventral cymbial tubercles, prolateral view (arrow points to ventral cymbial protuberance). 371. Embolus, prolateral view. 372. Radical division, retrolateral view (arrow points to basal spur). 373. Epigynum, ventral view. 374. Internal genitalia, ventral view. 375. Internal genitalia, dorsal view.