Cleotyche Emeljanov, 1997

Constant, Jérôme, Semeraro, Linda & Moir, Melinda L., 2022, Australian Cleotychini planthoppers: review of the genus Cleotyche Emeljanov, 1997 with three new species (Hemiptera: Fulgoromorpha: Dictyopharidae), European Journal of Taxonomy 836, pp. 66-95 : 69-70

Genus Cleotyche Emeljanov, 1997

Cleotyche Emeljanov, 1997: 78 [described, compared to Caliscelis De Laporte, 1833 , Parorgerioides de Bergevin, 1928 and Anorgeriopus Kusnezov, 1930 ].

Type species

Cleotyche mariae Emeljanov, 1997 , by monotypy and original designation.

Diagnosis

Only genus in the tribe.

Description

BODY. Pyriform; anterior part of body slightly compressed laterally.

HEAD. Trigons (triangular area delimited by carinae at anterior angles of vertex) well delimited. Frons subrectangular, narrow, parallel-sided, much longer than wide, with three longitudinal carinae and often two additional incomplete carinae on dorsal portion. Vertex subquadrate, longer than wide, with median carina. Clypeus elongate and narrow, subtriangular. Apical segment of the labium half as long as subapical segment.

THORAX. Pronotum smooth, elongate (slightly shorter than vertex along mid-line), tricarinate, with a single lateral keel on each side and posterior margin nearly straight. Mesonotum about ⅔ as long as pronotum, smooth with carinae nearly obsolete.

TEGMINA. Strongly reduced, without carina, truncate apically and slightly widening from base to apex; clavus not defined. Tegulae absent.

LEGS. Procoxae with triangular, lobe-shaped carina. Profemora and protibiae flattened and dilated, often foliate; with internal margin more or less evenly curved. Mesofemora and mesotibiae elongate and slender. Metatibiae with two lateral spines, one near tibiofemoral joint and one in distal half of tibia. Apex of metatibiae with 6 teeth separated in two groups: 2 teeth on internal side and 4 on external side. First and second segments of metatarsus with platellae instead of spines, except large lateral common teeth without setae. Arolium with one pair of chetoid sensilla. Claws with 3 setae.

ABDOMEN. Dorsoventrally flattened. Dorsum of abdomen without carinae.

MALE TERMINALIA. Pygofer narrow in lateral view, suboval and about much wider than high in posterior view; anterior and posterior margins rounded in lateral view; posterior margin deeply notched in dorsal and ventral view. Gonostyli rather compact, longer than high in lateral view, very elongate in ventral view; dorsal margin smoothly sinuate in lateral view; posteroventral margin rounded in lateral view; strong basidorsal lateral hook. Aedeagus elongate and narrow in dorsal view, with 2 pairs of membranous processes, each bearing a sclerotized spine; phallobase with 2 elongate sclerotized dorsal processes; connective elongate and narrow. Anal tube more or less oval in dorsal view with rounded apical margin; ventral margin sinuate; 2 strong apicolateral teeth directed ventrad.

Species included (type locality in parentheses)

Cleotyche (Griseotyche) blanda Emeljanov, 2011 (Stirling Range National Park, Western Australia) Cleotyche (Cleotyche) christinae sp. nov. (Cania Gorge National Park, Queensland)

Cleotyche (Cleotyche) francescoi sp. nov. (Eurimbula National Park, Queensland)

Cleotyche (Cleotyche) mariae Emeljanov, 1997 (Pearl Beach, New South Wales)

Cleotyche (Cleotyche) montana sp. nov. (Blackdown Tableland National Park, Queensland)

Identification key to the subgenera and species of Cleotychini Emeljanov, 1997

1. Ground colour pale grey ( Fig. 16A ); transverse white stripe on frons continuing laterally along body, becoming less distinct on abdomen ( Fig. 16B–C ); no white band along apical margin of tegmina ( Fig. 16A ) (subgenus Griseotyche ) ................................. C. (Griseotyche) blanda Emeljanov, 2011

– Ground colour dark brown ( Fig. 1 ); no transverse white stripe on frons nor lateral white stripe along side of body ( Fig. 2C ); white band along apical margin of tegmina ( Fig. 1 ) (subgenus Cleotyche ) 2

2. Profemora and protibiae narrower, more than 3 × as long as broad ( Fig. 1A, D ; Table 1 ) .............. 3

– Profemora and protibiae broader, less than 3 × as long as broad ( Fig. 1B–C ; Table 1 ) ................... 4

3. Profemora more slender, 3.78 × as long as broad; tibiae slightly broader than femora; vertex 1.56 × as long as broad ( Fig. 1A ) ............................................................. C. (Cleotyche) christinae sp. nov.

– Profemora broader, 3.16 × as long as broad; tibiae slightly narrower than femora; vertex 2.10 × as long as broad ( Fig. 1D ) .................................................................. C. (Cleotyche) montana sp. nov.

4. Profemora and protibiae broader, 2.55 and 2.73 × as long as broad, respectively; vertex less elongate, 1.71 × as long as than broad ( Fig. 1B ) ......................................... C. (Cleotyche) francescoi sp. nov.

– Profemora and protibiae narrower, 2.91 and 2.93 × as long as broad, respectively; vertex less elongate, 2.04 × as long as broad ( Fig. 1C ) ........................ C. (Cleotyche) mariae Emeljanov, 1997

Figures

Fig. 16. Cleotyche (Griseotyche) blanda Emeljanov, 2011, ♀, holotype (HMNH). A. Habitus dorsal view. B. Habitus, perpendicular view of frons. C. Habitus, lateral view. Photographs: © P. Kobor.

Fig. 1. Cleotyche (Cleotyche) Emeljanov, 1997 spp. of eastern Australia, ♂♂, dorsal view. A. C. (Cleotyche) christinae sp. nov. B. C. (Cleotyche) francescoi sp. nov. C. C. (Cleotyche) mariae Emeljanov, 1997. D. C. (Cleotyche) montana sp. nov.

Fig. 2. Cleotyche (Cleotyche) christinae sp. nov., ♂, holotype (QM). A. Habitus dorsal view.B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Head and thorax, dorsal view. G. Frons, perpendicular view. H. Head and thorax, lateral view. I. Distal portion of posterior leg, ventral view.