Pharaphodius rugosostriatus (Waterhouse, 1875)

Shabalin, S. A., 2018, To synonymy of Pharaphodius rugosostriatus (Waterhouse, 1875) (Coleoptera: Scarabaeidae), Far Eastern Entomologist 366, pp. 19-23 : 19-22

Pharaphodius rugosostriatus (Waterhouse, 1875)

Figs 1–7

Aphodius rugosostriatus Waterhouse, 1875: 92 . Type locality: "Kobé" [ Japan, Honshu ].

Aphodius (Pharaphodius) rugosostriatus: Schmidt, 1913: 124 ; Schmidt, 1922: 59; Nakane,

1960: 24; Balthasar, 1964: 68; Stebnicka, 1980: 231; Frolov, 1995: 292; Dellacasa &

Dellacasa, 2006: 135; Berlov & Shabalin, 2007: 15; Kim, 2012: 111; Akhmetova &

Frolov, 2014: 427.

Aphodius (Acrossus) rugosostriatus: Reitter, 1892(1891) : 247; d'Orbigny, 1896: 244.

Aphodius rugosostriatus: Miwa, 1930: 141 .

Pharaphodius rugosostriatus: Dellacasa et al., 2016a: 11 ; Dellacasa et al., 2016b: 146.

Aphodius (Pharaphodius) raddei Berlov, 1989: 392 . Type locality: Evseevka [ Russia:

Primorskii krai]. Synonymized by Stebnicka & Galante, 1991: 728.

Aphodius (Pseudacrossus) juxtus Petrovitz, 1972: 163 . Type locality: "Chemulpo" [ South

Korea, Incheon], syn. n.

lateral view, 3 – dorsal view, 4 – ventral view; 5 – labels.

Aphodius (Pseudacrossus) juxtus: Stebnicka, 1980: 205 ; Dellacasa & Dellacasa, 2006: 136;

Kim, 2012: 124.

Pseudacrossus juxtus: Dellacasa et al., 2016a: 11 ; Dellacasa et al., 2016b: 149.

TYPE MATERIAL. Holotype of Aphodius juxtus – ♂ ( Figs 1-5 ): " Korea / Chemulpo "

[white, handwritten] // "Aph. Pseudacross. / juxtus nov. spec. / Petrovitz" [red, printed] //

" TYPUS " [red, printed] // "Coll. / R. Petrovitz" [white, printed] // " MHNG / ENTO /

00013322" [white, printed]; Paratype of Aphodius juxtus – ♀ ( Figs 6, 7 ): " China / Prov .

Hupeh / Mts. Wu-schan" [white, printed] // "Aph. Pseudacross. / juxtus nov. spec. /

Petrovitz" [red, printed] // " PARATYPUS " [red, printed] // "Coll. / R. Petrovitz" [white,

printed] // " MHNG / ENTO / 00013323" [white, printed] // " A. rugosostriatus Waterh. / = A.

(Pseudacrossus) juxtus Petr. / syn. nov. / det. Z. Stebnicka 1989" [white, handwritten]. Type material deposited in Muséum d'histoire naturelle de Genève .

OTHER MATERIAL EXAMINED. Totally 969 specimens from Russia: Primorskii krai: 16–19.VII 1968, 1 specimen, M.A. Kascheev ; 67 km SW Vladivostok, Valley of

Ryazanovka River, 7.VIII 2009, 1 specimen, on light, S. Shabalin; the same locality, 8.VIII

2009, 3 specimens, in the cow excrements, S. Shabalin; the same locality, 16.VIII 2009, 2

specimens, on light, S. Shabalin; the same locality, 27.VIII 2009 1 specimen, on light, S.

Shabalin; the same locality, 29.IX 2017, 1 specimen, in the cow excrements, S. Shabalin; the same locality, 3. X 2017, 1 specimen, in the cow excrements, S. Shabalin; the same locality,

8.X 2017, 1 specimen, in the cow excrements, S. Shabalin; the same locality, 15.VII 2018, 32

specimens, in the cow excrements, S. Shabalin; the same locality, 22.VII 2018, 248 specimens, in the cow excrements, S. Shabalin; the same locality, 2.VIII 2018, 33 specimens, in the cow excrements, S. Shabalin; the same locality, 17.VIII 2018, 4 specimens, in the cow excrements, S. Shabalin; 49 km N Vladivostok, Razdolnoye village, 29.VIII 2018 1 specimen,

in the cow excrements, S. Shabalin; vicinity Ussuriisk, Utesnoe village, 21.VII 2018, 1 specimen, in the horse excrements, S. Shabalin; 20 km SE Ussuriisk, Dubovy Klyuch village,

5.VII 2018, 1 specimen, in the cow excrements, S. Shabalin; the same locality, 21.VII 2018,

37 specimens, in the cow excrements, S. Shabalin; the same locality, 1.VIII 2018, 87 specimens, in the cow excrements, S. Shabalin; the same locality, 13.VIII 2018, 1 specimen, in the cow excrements, S. Shabalin; 28 km SE Ussuriisk, Kaimanovka village, 5.VII 2018, 10

specimens, in the cow excrements, S. Shabalin; the same locality, 13.VII 2018, 16 specimens,

in the cow excrements, S. Shabalin; the same locality, 21.VII 2018, 3 specimens, in the cow excrements, S. Shabalin; the same locality, 1. VIII 2018, 4 specimens, in the cow excrements,

S. Shabalin; 16 km N Ussuriisk, Mikhailovka village, 21.VII 2018, 477 specimens, in the horse excrements, 94 km NE Ussuriisk, Knoring village, 17.VII 1981, 1 specimen, forest, on light, Kosheleva; Khabarovskii krai: 79 km SE Khabarovsk, Poletnoe village , on pastureland ,

9.VII 1968, 2 specimens, Radomskaya.

DISTRIBUTION. Russia (Primorskii krai, Khabarovskii krai), North Korea (Pyengannam-

do, Hwanghaebuk-do), South Korea (Gyeonggi-do, Chungcheongbuk-do, Gyeongsangbuk-

do, Jeollabuk-do, Jeollanam-do, Jeju-do), China (Heilongjiang, Jilin, Liaoning, Inner Mongolia,

Beijing, Tianjin, Hebei, Shandong, Shanxi, Ningxia, Gansu, Shaanxi, Henan, Jiangsu, Shang-

hai, Zhejiang, Anhui, Hubei, Sichuan, Taiwan, Fujian, Jiangxi, Hunan, Guizhou, Guangdong,

Guangxi, Yunnan, Hainan), Japan (Hokkaido, Honshu, Sado, Shikoku, Kyushu).

Figures

Figs 1–5. Holotype male of Aphodius juxtus: 1 – body, dorsal view; 2–4 – parameres: 2 –

Figs 6–7. Paratype female of Aphodius juxtus: 6 – body, dorsal view; 7 – labels.