Chlamydatus montanus Knight, 1964: 140

SCHUH, RANDALL T. & SCHWARTZ, MICHAEL D., 2005, Review of North American Chlamydatus Curtis Species, with New Synonymy and the Description of Two New Species (Heteroptera: Miridae: Phylinae), American Museum Novitates 3471 (1), pp. 1-56 : 37-44

Chlamydatus montanus Knight, 1964: 140 (n.sp.).

DIAGNOSIS: Recognized by the uniformly dark coloration of the dorsum, the reddish coloration of the legs, and the macropterous wing development. Distinguished from other species with unicolorous legs by the distinctive reddish quality in C. montanus , this condition otherwise occurring only in C. ruficornis , but that species easily distinguished by antennal segment 2 being red and weakly fusiform (fig. 2), whereas segment 2 black and of uniform diameter in C. montanus .

HOSTS: Apparently Artemisia spp. (Asteraceae) .

DISTRIBUTION: Western North America.

DISCUSSION: Knight (1964) described C. montanus from a single, somewhat mutilated specimen from Missoula, Montana, which is deposited in the U.S. National Museum. Kelton (1965) later recorded the species from Alberta and Saskatchewan, indicating the host as Pinus spp. Kelton’s host record is certainly in error, although his specimens undoubtedly represent a Chlamydatus species , based on his genitalic illustration ( Kelton, 1965). We collected specimens near Fort Collins, Colorado, probably sweeping an herbaceous Artemisia sp. and have a single specimen from near Kennedy Meadows in the Sierra Nevada mountains of California from Artemisia .

SPECIMENS EXAMINED: Holotype: female: ‘‘ Montana: Missoula, August 1, 1920, A. A. Nichol’ ’; deposited in the National Museum of Natural History , Washington, D.C.

Additional specimens: CANADA: Alberta: Canmore, August 27, 1952, L. A. Konotopetz, 1♀ (CNC). British Columbia : Chilcotin, Lee’s Corner, 5 mi E of, July 15, 1978, G. G. E. Scudder, 13 (UBC). Stewart Crossing, 16 km NW of, July 21, 1985, J. J. Robinson, 13 (UBC). Manitoba: Riding Mountain National Park, July 20, 1972, L. A. Kelton, 33 (CNC). Saskatchewan: Christopher Lake, July 15, 1959, A. R. and J. E. Brooks, 13 (CNC). Hudson Bay, August 24, 1954, A. R. Brooks, Wallis, 13, 1♀ (CNC). Yukon Territory: Jake’s Corner, August 24, 1981, C. S. Guppy, 1♀ (UBC). USA: California: Tulare Co. : 21 mi W of Rt 395 toward Kennedy Meadows, 1875 m, July 1, 1980, R. T. Schuh, Artemisia tridentata (Asteraceae) , 1♀ (AMNH). Colorado: Boulder Co.: Gold Hill, 8200 ft, August 6, 1973, J. Sawbridge, 13, 1♀ (OSU). Sunset, 8000 ft, July 19, 1903, E. P. Van Duzee, 13 (CNC). Larimer Co.: 46 mi W of Fort Collins, Fish Creek Picnic Grounds, Pingree Park Road, 7700 ft, July 14, 1986, R. T. Schuh and J. T. Polhemus, 53, 3♀ (AMNH).

Chlamydatus obliquus (Uhler)

Figures 1 , 3A , 5

Agalliastes obliquus Uhler, 1893: 378 (n.sp.).

DIAGNOSIS: Recognized by the relatively large size, total length 2.78–2.98, fore­ and middle femora pale, contrasting with dark hind femora, and the pronotum and hemelytra usually partially pale, as in the smaller C. schuhi , but less frequently dorsum entirely black. Dorsum clothed with shining, weakly woolly setae. Femoral coloration as in C. artemisiae , C. associatus , C. pallidicornis , and C. schuhi ; distinguished from those species by its larger size, the total length always at least 2.78 or more as opposed to a total length of 2.76 or less in the other three taxa; length of antennal segment 2 roughly equal to width of head in C. obliquus (75:74), whereas in C. associatus antennal segment 2 always longer than width of head (75:68) (table 1). Antennal segments 1 and 2 in males of C. pallidicornis usually entirely pale, whereas obliquus males always with antennal segments 1 and 2 black; vestiture in C. pallidicornis dark and neatly arranged, not shining and weakly woolly as in C. obliquus . Pale form of C. obliquus easily distinguished from C. schuhi by its larger size, total length in C. schuhi always being 2.66 or less. Vesica similar in form to several other North American spp. such as C. becki , C. brevicornis , and C. monilipes (figs. 4, 5).

HOSTS: Most commonly collected on Lupinus spp. (Fabaceae) and other herbaceous legumes growing on the forest floor.

DISTRIBUTION: Widely distributed in western North America.

DISCUSSION: We are designating a lectotype from material examined by Uhler in or­ der to assure stability of nomenclature for this taxon.

SPECIMENS EXAMINED: Lectotype: male: ‘‘Wasatch, Ut., 27­6.91, P. R. Uhler Collection, Agalliastes obliquus Uhler Utah, Agalliastes obliquus Uhler lectotype’’; deposited in the National Museum of Natural History, Washington, D.C.

Additional specimens: CANADA: Alberta: Coults, June 15, 1952, A. R. Brooks, L. A. Konotopetz, Lupinus argenteus (Fabaceae) , 173, 11♀ (CNC). Cypress Hills Provincial Park, Murray Hill Road, 4700 ft, July 18, 1990, M. D. Schwartz, 13 (AMNH). Frank, June 18, 1952, L. A. Konotopetz, 23 (CNC). Kananaskis Road, July 20, 1974, L. A. Kelton, 13 (CNC). Kananaskis Road, July 27, 1974, L. A. Kelton, 23, 2♀ (CNC). Lethbridge, July 13, 1938, R. W. Salt, Medicago sativa (Fabaceae) , 1♀ (CNC). Waterton Lakes National Park, July 22, 1923, J. Mc­ Dunnough, 3♀ (CNC). Waterton Lakes National Park, July 4–6, 26, 1972, L. A. Kelton, 183, 15♀ (CNC). British Columbia : Anarchist Mountain, Osoyoos, June 7, 1959, L. A. Kelton, R. Madge, Lupinus sp. (Fabaceae) , 103, 5♀ (CNC). Anarchist Mountain, Osoyoos, July 6, 1959, R. Madge, 13, 1♀ (CNC). Anarchist Mountain, Osoyoos, July 30, 1959, L. A. Kelton, 13 (CNC). Anarchist Mountain, Osoyoos, July 13, 1970, L. A. Kelton, 253, 1♀ (CNC). Aspen Grove, July 10, 1975, L. A. Kelton, 63 (CNC). Christina Lake, July 12, 1970, L. A. Kelton, 13 (CNC). Cranbrook, July 23, 1959, L. A. Kelton, 13, 4♀ (CNC). Creston, July 22, 1959, R. Madge, 1♀ (CNC). Crowsnest Provincial Park, 4 km W of Crows Nest Pass, 1396 m, September 1, 1993, M. D. Schwartz, Lupinus argenteus (Fabaceae) , 13 (AMNH). Elko, E Kootenay, July 9, 1949, H. B. Leech, 13 (CAS). Fairview, White Lake , July 7, 1996, G. G. E. Scudder, 6♀ (UBC). Grand Forks, June 13, 1950, H. R. MacCarthy, 13 (CNC). Jaffray, July 17, 1974, L. A. Kelton, 13 (CNC). Kaleden, July 2, 1975, L. A. Kelton, 23 (CNC). Madden Lake, Oliver, July 8, 1959, L. A. Kelton, Rhus sp. (Anacardiaceae) , 13, 1♀ (CNC). Manning Provincial Park, Blackwall, July 23, 1970, L. A. Kelton, 23 (CNC). McIntyre Creek, Oliver, May 27, 1959, R. E. Leech, Balsamorhiza sp. (Asteraceae) , 13 (CNC). McIntyre Creek, Oliver, July 4, 1959, L. A. Kelton, 13 (CNC). Merritt, 10 mi S of, July 19, 1959, L. A. Kelton, 33, 4♀ (CNC). 10 mi W of Princeton, July 31, 1957, N. H. Anderson, 2♀ (CNC). Merritt, 8 mi SE of, July 19, 1970, P. Oman, 33 (OSU). Nicola, July 10, 1932, G. J. Spencer, 1♀ (UBC). Nicola, June 14, 1963, G. G. E. Scudder, 33, 4♀ (UBC). Okanangan Valley, June 1, 1996, R. G. Foottit, D. R. Gillespie ( Fabaceae ) (CNC). Oliver, May 25, 1959, R. Madge, Astragalus sp. (Fabaceae) , 13, 1♀ (CNC). Oliver, May 27, 1959, L. A. Kelton, 13 (CNC). Oliver, May 29, 1959, R. E. Leech, E. E. McDougall, Lupinus sp. (Fabaceae) , 473, 25♀ (CNC). Oliver, June 20, 1975, L. A. Kelton, 13 (CNC). Oliver, July 2, 1975, L. A. Kelton, 13 (CNC). Oliver, July 19, 1970, L. A. Kelton, 13 (CNC). Osoyoos, June 24, 25, 1953, J. R. McGillis, 83, 34♀ (CNC). Osoyoos, 900 m, July 5, 1996, R. G. Foottit, D. R. Gillespie, 1♀ (CNC). Princeton, 870 m, July 4, 1996, R. G. Foottit, D. R. Gillespie (CNC). Princeton, June 4, 1961, G. G. E. Scudder, 33 (UBC). Quilchena Pennask Lake Rd, km 7.1, June 29, 2001, G. G. E. Scudder, 33, 5♀ (UBC). Richter Pass, June 6, 1959, E. E. McDougall, 13 (CNC). Richter Pass, Osoyoos, May 24, 1959, L. A. Kelton, Lupinus sp. (Fabaceae) , 463, 25♀ (CNC). Rock Creek, June 7, 1959, L. A. Kelton and R. E. Leech, 223, 8♀ (CNC). Ross Lake, Okanagan Falls, June 5, 1959, R. Madge, 13 (CNC). Sawmill Lake, Oliver, June 30, 1959, L. A. Kelton, Lupinus sp. (Fabaceae) , 2♀ (CNC). Summerland, July 10, 1975, L. A. Kelton, 13 (CNC). Summit Lake, mi 392 Alaska Highway, 4500 ft, July 8, 1959, R. E. Leech, 13 (CNC). Vaseaux Lake, Oliver, May 23, 1959, R. E. Leech, 1♀ (CNC). Vaseaux Lake, Oliver, May 29, 1959, E. E. MacDougall, 33, 3♀ (CNC). Vaseaux Lake, Oliver, June 26, 1959, L. A. Kelton, 33, 5♀ (CNC). Vernon, June 26, 1949, C. L. Neilson, 33 (CNC). Vernon, June 30, 1940, R. L. Usinger, 13 (UCB). White Lake, Oliver , May 23, 27, 1959, E. E. MacDougall, 53, 9♀ (CNC). White Lake, Oliver , May 28, June 10, 29, 1959, L. A. Kelton, Lupinus sp. (Fabaceae) , 263, 23♀ (CNC). Wycliffe, June 8, 1961, G. G. E. Scudder, 13 (UBC). USA: Arizona: Apache Co. : 4.8 mi S of Hwy 60 on FR 117, 7930 ft, September 19, 1998, J. C. Schaffner, 23 (TAMU). Coconino Co.: Grand Canyon, south Rim, June 24, 1930, R. L. Usinger, 43 (UCB). Jacob Lake, June 5, 1969, W. F. Chamberlain, 53 (TAMU). Greenlee Co.: 8 mi S of Hannagan Meadow, Blue Vista scenic viewpoint, September 19, 1998, J. C. Schaffner, 13 (TAMU). California: Alpine Co. : Markleeville, SE of on Rt 89, 1.5– 5 mi NW jct. of Rts 4/89, July 5, 1994, Schwartz, Lupinus sp. (Fabaceae) , 63 (AMNH). Lassen Co.: Susanville, June 17, 1959, L. A. Kelton, R. Madge, Lupinus sp. (Fabaceae) , 263, 21♀ (CNC). Modoc Co.: 2.5 mi S of Rt 139 towards Lookout, 1440 m, July 6, 1979, R. T. and J. Schuh, Lupinus sp. (Fabaceae) , 1403 (AMNH). 8 mi S of Rt 139 towards Lookout, 1420 m, July 6, 1979, R. T. and J. Schuh, Lupinus sp. (Fabaceae) , 43 (AMNH). 18 mi S of Rt 139 towards Lookout, 1420 m, July 6, 1979, R. T. and J. Schuh, Lupinus sp. (Fabaceae) , 73 (AMNH). 5 mi S of Inspection Station, Highway 139, July 17, 1955, Joe Schuh, 33 (OSU). Shasta Co.: 8 mi W of Fall River Mills, 845 m, July 7, 1979, R. T. and J. Schuh, Iva axillaris (Asteraceae) , 23 (AMNH). Mount Shasta Ski Area, 8000 ft, August 31, 1979, J. D. Lattin, Lupinus sp. (Fabaceae) , 13 (OSU). Siskiyou Co.: 1 mi W of Bartle, 1220 m, July 7, 1979, R. T. and J. Schuh, 13 (AMNH). 12.3 mi N of St. Hwy. 89 on Powder Hill Road, July 19, 1985, G. M. Stonedahl and J. D. McIver, Chrysothamnus viscidiflorus (Asteraceae) , 13 (AMNH). Big Flat, August 4, 1931, R. L. Usinger, 33 (UCB). McCloud, July 5, 1952, M. Cazier, W. Gertsch, and R. Schrammel, 83 (AMNH). Colorado: Garfield Co.: Baxter Pass (Radio Tower), 14 mi SE of CO St. Border, 8200 ft, July 9, 1981, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 13 (AMNH). Gunnison Co.: Taylor Park, July 25, 1982, J. T. Polhemus, 203 (JTP). Jackson Co.: Cowdrey, 8000 ft, August 15, 1968, L. A. Kelton, Chrysothamnus sp. (Asteraceae) , 13, 1♀ (CNC). Near Cowdrey, Colorado–Wyoming state line on Rt 125, August 4, 1979, J. T. Polhemus, 13 (JTP). Montezuma Co. : Mesa Verde National Park, July 13, 1930, R. L. Usinger, 143 (UCB). Montrose Co.: 27 mi E of Nucla, 9200 ft, August 14, 1987, T. J. Henry, 13, 13♀ (USNM). Routt Co.: Steamboat Springs, 6700 ft, July 11, 1964, H. H. Knight, 43, 1♀ (AMNH, CNC). Steamboat Springs, July 15, 1942, 13 (AMNH). Steamboat Springs, July 23, 1983, J. T. Polhemus, 63 (JTP). Idaho: Bear Lake Co.: Montpelier, 6100 ft, July 6, 1920, 13 (AMNH). Bonner Co.: Pend Orielle River, Laclede, July 4, 1966, W. Gagne, J. Haddock, 43 (UCB). Fremont Co.: 20 mi N of Ashton, Mac’s Inn, July 29, 1982, S. E. Cummings, 13 (UNHP). Kootenai Co.: Post Falls, July 13, 1929, R. A. Flock, 13 (UCR). Latah Co.: 6.4 mi S of Troy on Rt 99, June 8, 1989, R. S. Zack, 23, 2♀ (WSU). Moscow, May 17, 1934, G. W. Dahmen, 1♀ (CNC). Moscow Mountain, July 31, 1972, L. A. Kelton, 63, 6♀ (CNC). Lemhi Co.: Bannock Pass, on Rt 29, Beaverhead Mountains, Salmon National Forest, July 31, 1994, M. D. Schwartz, Lupinus argenteus (Fabaceae) , 13 (CNC). Nez Perce Co.: Lewiston, 550 ft, May 30, 1932, T. A. Brindley (CNC). Lewiston, 550 ft, May 30, 1932, T. A. Brindley, 43 (AMNH, CNC, USNM). Oneida Co. : Salyer Cow Camp, July 23, 1971, W. J. Hanson, 13 (USU). Montana: Beaverhead Co.: 5 mi E of Wise River, July 21, 1983, J. D. Pinto, 23 (UCR). Clark Canyon Dam, July 14, 1982, S. E. Cummings, 13 (UNHP). Flathead Co.: Glacier National Park, July 28, 1972, L. A. Kelton, 33, 1♀ (CNC). Gallatin Co.: Bozeman, 12 mi E of, July 23, 1983, J. D. Pinto, 53 (UCR). Moose Flat Campground, 26 mi S of Bozeman Hot Springs, on Rt 191, 5700 ft, August 10, 1986, Schuh, Schwartz, Stonedahl, 13 (AMNH). Elkhorn Ranch, Gallatin River, Upper Gallatin Canyon, 7000 ft, August 1, 1928, J. McDunnough 13, 7♀ (CNC). Madison Co.: 15 mi SE of Virginia City, July 12, 1982, S. E. Cummings, 13 (UNHP). Powell Co.: 1.4 mi SW of Ovado on Rt 200, May 28, 1992, R. S. Zack, 23, 2♀ (WSU). Silverbow Co.: Butte, July 20, 1983, J. D. and D. R. Pinto, 113 (UCR). Nevada: Carson City Co. : Carson City, June 25, 1929, R. L. Usinger, 53 (CAS). Elko Co.: Ruby Mts., Lomoille Canyon, E of Powerhouse Picnic Area, 6200 ft, June 16, 1983, R. T. Schuh and M. D. Schwartz, Artemisia dracunculus (Asteraceae) , 113 (AMNH). Humboldt Co.: 4 mi E of Winnemucca, June 23, 1971, P. Oman, 13 (OSU). Lander Co.: 6 mi E of Austin, July 2, 1961, H. and A. Howden, 1♀ (CNC). Nye Co.: Atomic Test Site, 2 mi W of Tippapah Hwy on Mine Mt. Road, 4400 ft, June 7, 1983, Schuh, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 23 (AMNH). Atomic Test Site, 1.5 mi W Area 12 camp on Stockade Wash Rd, June 7, 1983, Schuh, Schwartz, Stonedahl, Lupinus argenteus (Fabaceae) , 23 (AMNH). Manhattan, 3.5 mi SE of, Toiyabe National Forest, 7000 ft, July 13, 1980, R. T. Schuh and G. M. Stonedahl, Lupinus sp. (Fabaceae) , 13 (AMNH). Washoe Co.: Nixon, June 29, 1927, E. P. Van Duzee, 13 (CAS). New Mexico: Colfax Co. : 5 mi W of Cimmaron, June 16, 1992, G. M. Chamberlain, 13 (TAMU). Oregon: Baker Co.: 3.3 mi E of Halfway, June 7, 1970, P. Oman, 13 (OSU). Crook Co.: 10 mi E of western boundary of Ochoco National Forest on Rt 26, R19E T13S, sec. 3, June 22, 1979, R. T. Schuh, 33 (AMNH). Deschutes Co.: Brothers, 8 mi SE of, July 9, 1968, J. D. Lattin, 13 (OSU). Millican, 1 mi S of, 4380 ft, June 21, 1979, G. M. Cooper, 13 (OSU). Millican, 14 mi S of, R15E, T20S, sec. 34, 5400 ft, July 21, 1979, M. D. Schwartz and R. T. Schuh, 103 (AMNH). Pine Mountain, R15E, T21S, Section 10, 5400 ft, June 21, 1979, P. Oman, 13 (OSU). Grant Co. : 16 mi NE of Prairie City, July 13, 1991, W. F. Chamberlain, 23 (TAMU). Harney Co.: Burns, 15 mi W of, on State Rd 20, rest stop, 4850 ft, June 14, 1981, M. D. Schwartz, Agropyron sp. (Poaceae) , 13 (AMNH). Fish Lake, Steens Mts., 7200 ft, August 12, 1958, K. M. Fender, 233 (OSU). Frenchglen, 8 mi E of, July 16, 1957, J. D. Lattin, Lupinus caudatus (Fabaceae) , 23 (OSU). Frenchglen, 13 mi E of, 6500 ft, August 13, 1971, P. Oman, 13 (OSU). Lily Lake, 13 mi E of Frenchglen, 7200 ft, July 10, 1968, P. Oman, 23 (OSU). Malheur Lake, August 17, 1958, K. M. Fender, 13 (OSU). T36S, R35E, sec. 8 N1/2, dunes, August 7, 1979, Lightfoot and Cobb, 13 (OSU). Jackson Co.: 0.5 mi E of Pinehurst, 3550 ft, June 27, 1979, M. D. Schwartz, 83 (AMNH). Just E of Pinehurst, 1340 m, June 27, 1979, R. T. and J. Schuh, Lupinus sp. (Fabaceae) , 83 (AMNH). Klamath Co.: between Worden and Keno, 1280 m, June 27, 1979, R. T. and J. Schuh, 33 (AMNH). Chiloquin, Rt 97, July 4, 1982, G. M. Stonedahl and T. J. Henry, Lupinus sp. (Fabaceae) , 223 (AMNH). Skookum Meadow, R9E, T27S, Section 16, July 17, 1979, P. Oman, 13 (OSU). Lake Co.: Hart Mountain, June 17, 1938, Gray and Schuh, 13 (AMNH). Linn Co.: Tombstone Prairie, 35 mi E of Sweet Home, 4500 ft, August 15, 1970, J. D. Lattin, 13 (OSU). Morrow Co.: Boardman, May 11, 1960, K. Goeden, 13 (OSU). Umatilla Co.: 0.4 mi S of Jct of Rts 244 and 395, Lupinus sp. (Fabaceae) , 153 (AMNH). Near Ukiah, June 23, 1935, Joe Schuh, 33, 2♀ (AMNH, CNC). Union Co.: Imbler, 3 mi NW of, July 6, 1956, J. D. Lattin, 13 (OSU). Woodland Campground, 4.5 mi E of Tollgate, 5000 ft, August 4, 1986, Schuh, Schwartz, Stonedahl, Potentilla fruticosa (Rosaceae) , 13 (AMNH). Wallowa Co.: Enterprise, June 4, 1924, 13 (OSU). Enterprise, 30 mi N of, July 30, 1968, P. Oman, 13 (OSU). Joseph, June 6, 1970, P. Oman, 73 (OSU). Joseph, June 13, 1973, Oman and Musgrave, 13 (OSU). Joseph, July 10, 1969, P. Oman, 253 (OSU). Joseph, 15 mi NE of, June 19, 1968, P. Oman, 13 (OSU). Joseph, 24 mi E of, July 11, 1969, P. Oman, 13 (OSU). Joseph, 35 mi N of, June 21, 1968, P. Oman, 233 (OSU). Lostine, 3.5 mi S of, July 21, 1970, P. Oman, 33 (OSU). Wasco Co.: Maupin, June 5, 1962, K. M. and D. M. Fender, 53 (OSU). The Dalles, June 3, 1970, P. Oman, 13 (OSU). Tyghe Valley, 6 mi S of, June 6, 1962, K. M. Fender, 13 (OSU). Wheeler Co.: 4 mi SW of Fossil, June 5, 1979, P. Oman, Balsamorhiza sp. (Asteraceae) , 13 (AMNH). Utah: Bear Lake Co.: Garden City, July 6, 1968, G. R. Knowlton, H. H. Knight, 13 (CNC). Box Elder Co. : Blue Creek, June 24, 1932, G. F. Knowlton, 13 (USU). Cache Co.: 14 mi S on FS Rd 055 off UT St. Rt 89, T13N, R4E S15, July 25, 1981, Schwartz, Artemisia tridentata (Asteraceae) (AMNH). Garfield Co.: 6 mi W of Starr Springs, Mount Hillers, 6700 ft, June 20, 1983, Schuh and Schwartz, Lupinus sp. (Fabaceae) , 103 (AMNH). San Juan Co.: La Sal Mts., nr. Brumley Creek, T27S, R24E, 8000 ft, June 11, 1982, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 33 (AMNH). Sevier Co.: Clear Creek Narrows Summit on Rt 4, 4224 m, July 15, 1980, R. T. Schuh and G. M. Stonedahl, 93 (AMNH). Uintah Co.: Oaks Park, August 2, 1965, G. F. Knowlton, 143 (USU). Utah Co.: Provo City, April 15, 1965, A. T. Whitehead, 13 (UCB). Washington: Adams Co.: 17 mi W of Washtucna, Sand Hills Park, June 13, 1989, R. S. Zack, 13 (WSU). Asotin Co.: 2.5 mi S of Anatone, 2 mi N Rattlesnake Summit, 3900 ft, August 4, 1986, Schwartz, Schuh, Stonedahl, Lupinus sp. (Fabaceae) , 13 (AMNH). 3 mi N of Anatone, June 5, 1970, P. Oman, 13 (OSU). Benton Co.: Hanford Site ALE, near ALE HQ, T11N, R26E, sec. 34, 1200 ft, May 4, 1974, R. S. Zack, 23 (CNC). Hanford Site ALE, T11N, R26E, 1200 ft, May 4, 1994 and June 1, 1994, R. S. Zack, 83, 8♀ (WSU). Douglas Co.: 14 mi N of Wenatchee, Columbia View Orchards , June 20, 1985, R. J. Herman, 63, 6♀ (WSU). Ferry Co.: Republic, August 6, 1931, H. T. Peters, 13 (KU). Garfield Co.: Colton, July 6, 1948, C. C. Shelton, 103, 10♀ (WSU). Kittitas Co.: Cliffdell, July 7, 1935, R. H. Beamer, 13 (KU). Klickitat Co.: Bickleton, June 15, 1935, W. Downes, 13 (UBC). Okanogan Co.: Okanogan, 11 mi W of, May 19, 1973, P. Oman, 13 (OSU). Sidley Lake, Molson, July 7, 1964, G. G. E. Scud­ der, 43, 6♀ (CNC). Pierce Co.: Mount Rainier, July 6, 1935, R. H. Beamer, 13 (KU). Whitman Co.: Smoot Hill, 2 mi N of Albion, July 9, 1989, R. S. Zack, 13, 1♀ (WSU). Steptoe Butte, 4 mi NE of Steptoe, June 11, 1989, R. S. Zack, 43, 4♀ (WSU). Yakima Co.: 13 mi SW of Ellensburg, T16N, R17E, sec. 6, June 13, 1989, R. S. Zack, 13 (WSU). Wyoming: Big Horn Co.: 24 mi W of Burgess Jct on road to radar station, 9000– 9200 ft, August 12, 1986, Schuh, Schwartz, Stonedahl, Lupinus argenteus (Fabaceae) , 43 (AMNH). Big Horn Mts., 6500 ft, August 17, 1927, H. H. Knight, 13 (USNM). Carbon Co.: Saratoga, July 29, 1931, H. H. Knight, 13 (USNM). Fremont Co.: Wind River Mts., Shoshone National Forest, Popoagie Campground on Rt 131, August 14, 1986, Schwartz and Stonedahl, Shepherdia canadensis (Elaeagnaceae) , 23 (AMNH). Lincoln Co.: 10 mi E of Alpine near Wolf Creek Campground, July 21, 1981, M. D. Schwartz, Lupinus sp. (Fabaceae) , 313 (AMNH). Shoshone Co.: 19 mi E of Cooke City on Rt 212, 8000 ft, August 11, 1986, Schuh, Schwartz, Stonedahl, Lupinus sp. (Fabaceae) , 33 (AMNH). Teton Co.: Grand Teton National Forest, August 7, 1972, L. A. Kelton, 13 (CNC). Grand Teton National Park, 2 mi SE Colter Bay Village, 7000 ft, July 22, 1981, M. D. Schwartz, 23 (AMNH).

Chlamydatus opacus (Zetterstedt)

Figures 1 , 3G , 5

Capsus opacus Zetterstedt, 1838: 279 (n.sp.).

DIAGNOSIS: Easily recognized by the brachypterous condition of the hemelytra, the membrane small, but present, and the brown and cream coloration (fig. 1). Most easily confused with C. californicus in North America, but hemelytra truncate in that species, staphylinoid in form, and the membrane completely lacking.

HOSTS: No host records exist for this species.

DISTRIBUTION: Holarctic. Occurring only at very high latitudes in the New World.

DISCUSSION: All Palearctic specimens we examined lacked the hemelytral membrane.

SPECIMENS EXAMINED: AUSTRIA: Neusiedl , July 21, 1960, G. G. E. Scudder, 1♀ ( CNC) . Neusiedler Lake , July 21, 1960, G. G. E. Scudder, 13 ( CNC) . CANADA: Manitoba: Churchill , July 12, 1952, J. G. Chillcott, 3♀ ( CNC) . Northwest Territories: Geillini Lake , August 14, 1952, J. G. Chillcott, 13, 5♀ ( CNC) . Quebec: Great Whale River , July 12 and 18, 1949, J. R. Vockeroth, 213, 11♀ ( CNC) . GERMANY: Holstein, Plön , July 26, 1964, G. G. E. Scudder, 33, 5♀ ( CNC) . RUSSIA: Khabarovak, E. Siberia , August 26, 1979, G. G. E. Scudder, 13, 1♀ ( CNC) . Khekhtzir Forest , nr Khabarovak, E. Siberia, August 26, 1979, G. G. E. Scudder, 13, 1♀ ( CNC) . Moscow Hills , August 5, 1968, G. G. E. Scudder, 43 ( CNC) .

Chlamydatus pallidicornis Knight

Figures 1 , 3A , 5

Chlamydatus pallidicornis Knight, 1964: 138 (n.sp., host).

DIAGNOSIS: Recognized by the uniformly dark coloration of the dorsum, the pale whitish or yellowish fore­ and middle femora and contrastingly dark hind femora, the dorsal vestiture composed of simple, dark, neatly arranged, nonshining setae, and by the usually pale antennal segments 1 and 2 (fig. 2). Most similar in overall coloration to C. artemisiae , C. associatus , and C. obscurus . Distinguished from C. artemisiae , C. associatus , and C. obliquus by the dorsal vestiture being shining and somewhat disordered in those species, whereas dorsal vestiture dark and neatly arranged in C. pallidicornis . Antennal segment 2 shorter than width of head in C. pallidicornis (59:70), similar to C. artemisiae , whereas antennal segment 2 about equal to width of head in C. obliquus (75:74), and longer than the width of the head in C. associatus (75:68) (table 1); in males of C. artemisiae , C. associatus , and C. obliquus antennal segments 1 and 2 dark colored, or mostly so, whereas segment 2 usually pale in males of C. pallidicornis . Easily distinguished from C. obliquus specimens with a totally dark dorsum by total length in that species always being at least 2.78, whereas total length in C. pallidicornis 2.54 or less.

HOST: Potentilla plattensis ( Knight, 1964) ; Potentilla sp. (Rosaceae) , on the basis of the only available specimen label data.

DISTRIBUTION: Western Canada and higher latitudes in the western United States.

DISCUSSION: Knight (1964) recorded the host of this species as Potentilla plattensis , with every indication that the bugs were breeding on the plant. Host information available to us is much less clear­cut, suggesting the need for further fieldwork.

SPECIMENS EXAMINED: Holotype: male: ‘‘ Shoshone N. Forest, Wyo., Aug. 14, 1927, H. H. Knight’ ’; deposited in the National Museum of Natural History , Washington, D.C.

Additional specimens: CANADA: Alberta: Cypress Hills Provincial Park, Top Road, 9.6 km E of Rt 41 jct, July 15, 1990, M. D. Schwartz, 1♀ (CNC). Elkwater Park, July 12, 1952 and August 15, 1952, A. R. Brooks, L. A. Konotopetz, 2♀ (CNC). British Columbia : Anarchist Mountain, Osoyoos, July 13, 1970, L. A. Kelton, 33, 2♀ (CNC). Blackwall, Manning Provincial Park, July 23, 1970, L. A. Kelton, 13 (CNC). Christina Lake, July 12, 1970, L. A. Kelton, Potentilla sp. (Rosaceae) , 183, 24♀ (CNC). Merritt, 10 mi S of, July 19, 1959, L. A. Kelton, 13, 4♀ (CNC). Merritt, 12 mi S of, July 19, 1959, L. A. Kelton, 23, 2♀ (CNC). USA: Arizona: Apache Co. : Big Lake, Apache National Forest, 9100 ft, July 11, 1968, August 14, 1967, L. A. Kelton, 463, 69♀ (CNC). Colorado: Archuleta Co.: Pagosa Springs, July 22, 1968, L. A. Kelton, 83, 2♀ (CNC). Jackson Co.: Muddy Pass, Routt National Forest, 8600 ft, August 21, 1968, L. A. Kelton, 63, 5♀ (CNC). Rabbit Ears Pass, 9400 ft, July 4, 1978, J. T. Polhemus, 1♀ (JTP). Rabbit Ears Pass, August 3, 1947, L. M. and R. H. Beamer, 53, 2♀ (KU). La Platta Co.: La Platta, San Juan National Forest, 8500 ft, July 19, 1968, L. A. Kelton, 13, 1♀ (CNC). Routt Co.: Steamboat Springs, 6900 ft, July 15, 1964, H. H. Knight, 1♀ (CNC). San Juan Co.: near Ophir, 8500 ft, July 7, 1980, J. T. Polhemus, 83, 4♀ (JTP). Idaho: Blaine Co.: Ketchum, July 31, 1955 and August 1, 1955, B. P. Bliven, 23 (CAS). Idaho Co.: Lowell, Clearwater National Forest, July 30, 1972, L. A. Kelton, 1♀ (CNC). New Mexico: Otero Co.: Cloudcroft, June 27, 1940, R. H. Beamer, 3♀ (KU). Oregon: Crook Co.: 10 mi E of W boundary of Ochoco National Forest on Rt 26, T13S, R19E, sec. 3, June 22, 1979, R. T. Schuh, 13 (AMNH). Ochoco National Forest, T14S, R18E, sec. 3, June 22, 1979, M. D. Schwartz, Senecio sp. (Asteraceae) , 13, 1♀ (OSU). Lake Co.: Hart Mountain Refuge, Hot Spring Campground, July 19, 1971, J. D. Lattin, 1♀ (OSU). Union Co.: 4.5 mi E of Tollgate, Woodland Campground, 5000 ft, August 4, 1986, Schuh, Schwartz, Stonedahl, 13, 4♀ (AMNH). Wallowa Co.: 15 mi N of Enterprise, August 18, 1969, P. W. Oman, 23, 2♀ (OSU). 8 mi E of Joseph, July 10, 1969, P. W. Oman, 13 (OSU). Wheeler Co.: 4.5 mi S of Mitchell on Summit Prairie Road, June 22, 1979, M. D. Schwartz, Cercocarpus ledifolius (Rosaceae) , 33 (OSU). Utah: Cache Co.: Ant Valley, July 6, 1973, G. E. Bohart and G. F. Knowlton, 13, 1♀ (USU). Blacksmith Fork Canyon, June 28, 1971, G. F. Knowlton, 13 (USU). Franklin Basin, July 4, 1972, G. E. Bohart, 53 (USU). Logan, July 17, 1973, G. F. Knowlton, 23, 1♀ (UCD). Richmond, August 2, 1971, G. F. Knowlton, 1♀ (USU). Rock Creek, July 22, 1976, W. J. Hanson, 1♀ (USU). Washington Co.: Zion National Park, August 18, 1929, P. W. Oman, 13 (KU). Washington: Benton Co.: Hanford Site ALE, radio telescope, June 9, 1994, R. S. Zack, 33, 3♀ (WSU). Columbia Co. : Dayton, July 12, 1937, R. E. Miller, 1♀ (CNC). Okanogan Co.: 8 mi WNW of Republic, Sweet Creek, 3600 ft, July 20, 1978, N. Herman, 103, 7♀ (AMNH). Whitman Co.: Steptoe Butte State Park, 4 mi NE of Steptoe, June 11, 1989, R. S. Zack, 23 (WSU). Near Colton, Palouse Prairie, Kramer Strip, June 25, 1983, W. J. Turner, 63, 2♀ (WSU). Wyoming: Fremont Co.: Shoshone National Forest, August 14, 1927, H. H. Knight, 23, 1♀ (CNC). Sublette Co.: 25 mi SE of Jackson, Hobart River, June 27, 1966, W. Gagne and J. Haddock, 13 (UCB). Teton Co.: Grand Teton National Forest, August 7, 1972, L. A. Kelton, 13, 3♀ (CNC). Grand Teton National Park, near Kelly, July 24, 1982, S. C. Williams, 2♀ (CAS).

Chlamydatus pulicarius (Fallén)

Figures 2 , 3B, C , 5 , 9

Lygaeus pulicarius Fallén, 1807: 95 (n.sp.).

Chlamydatus arcuatus Knight, 1964: 139 (n.sp.). NEW SYNONYMY.

Chlamydatus auratus Kelton, 1965: 1133 (n.sp., host). NEW SYNONYMY.

DIAGNOSIS: Recognized by its relatively large size, total length 2.51–3.09, the often ovate to obovate body form, and the unicolorous hemelytra being submacropterous in form and just covering the abdomen (fig. 2), or more rarely macropterous (fig. 2) and being elongate and nearly parallel sided. Femora ranging from almost entirely yellowish to almost entirely dark (fig. 3B,C). The coloration of antennal segments 1 and 2 varying from completely pale to almost entirely black. Most easily confused with C. keltoni and C. pullus on the basis of frequent submacroptery in all of these species and conformation of hemelytra to abdomen, but total length in those species 2.61 or less, whereas C. pulicarius always 2.51 or more, and male genitalia (fig. 5) more highly convoluted and vesica with a longer apical process in C. pulicarius than in C. keltoni (fig. 4); variation in coloration of appendages in C. pullus similar to that found in C. pulicarius . Possibly also confused with C. uniformis on the basis of size, but that species jet black with all legs entirely black and occurring in southern California and adjacent northern Mexico, whereas C. pulicarius with at least tibiae always pale (figs. 2, 3B,C) and restricted to much higher latitudes or higher altitudes.

HOSTS: Substantial numbers of specimens have been collected from the following plant taxa: Artemisia sp. , Hieracium sp. (Asteraceae) , Stellaria longifolia (Caryophyllaceae) , Lathyrus sp. , Vicia sp. (Fabaceae) , Ranunculus sp. (Ranunculaceae) , and Potentilla argentea (Rosaceae) . Other taxa may serve as breeding hosts, but the available specimen data are not sufficient to draw that conclusion at the present time.

DISTRIBUTION: Holarctic. Occurring primarily at higher latitudes and higher altitudes in North America, but extending as far south as West Virginia in the eastern United States.

DISCUSSION: Knight (1964) described C. arcuatus as widely distributed in the Rocky Mountain system and Kelton (1965) described C. auratus as occurring across Canada and in adjacent New York. The characters used by Knight and Kelton to distinguish both of these nominal species from C. pulicarius , particularly the vestiture, are variable and difficult to interpret. The genitalia of both appear identical to C. pulicarius , and after having examined the holotypes and substantial other material of C. arcuatus and C. auratus , we have concluded that both are synonymous with C. pulicarius new synonymy.

Kelton (1965) had a narrowly conceived diagnosis of this species, largely based on color. He assigned to C. pulicarius only specimens from Alaska, Alberta, and British Columbia . Kelton (1980) indicated that he doubted that C. pulicarius actually occurred in North America. Wheeler and Henry (1992) indicated that they had no definitive records of the species from North America and therefore excluded it from their treatment of Holaractic taxa. Our conception of the species is based on the distinctive genitalic structure, an approach that construes the taxon as being much more widely distributed than was thought to be the case by all previous authors. We have not examined material originally studied by Fallén, but rather have relied on the characterizations of authors such as Kerzhner (e.g., 1964, in litt.) in forming our understanding of the correct identity of this taxon.

SPECIMENS EXAMINED: Holotype: Chlamydatus arcuatus : male: ‘‘Wolf Creek Pass, Colo., Aug. 12, 1925, H. H. Knight’’; deposited in the National Museum of Natural History, Washington, D.C. Holotype: Chlamydatus auratus : male: ‘‘One Sided Lake, ONT., 25.VI.1960, Kelton & Whitney, Type No. 8901’’; deposited in the Canadian National Collection of Insects, Ottawa.

Additional specimens: AUSTRIA: Igls, Tirol, 900 m, September 17, 1953, J. R. Vockeroth, 13 (CNC). Neusiedl, July 21, 1960, G. G. E. Scudder, 1♀ (CNC). Obergurgl, Tirol, 900 m, August 12, 1953, J. R. Vockeroth, 2♀ (CNC). Ofztal, Tirol, 700 m, July 2, 1953, J. R. Vockeroth, 1♀ (CNC). Schönfeld, August 21, 1960, G. G. E. Scudder, 43, 1♀ (CNC). CANADA: Alberta: Athabasca, Peace River, July 1, 1900, Raup, 13 (AMNH). Coal Valley, August 31, 1970, L. A. Kelton, 23, 6♀ (CNC). Cypress Hills Provincial Park, Murray Hill Road, July 18, 1990, M. D. Schwartz, 1♀ (CNC). Drumheller, June 18, 1957, Brooks, MacNay, C. auratus paratypes, 13, 2♀ (CNC). Edmonton, August 5, 1952, L. A. Konotopetz, 13, 1♀ (CNC). Elkwater Park, July 7, 12, 23, 30, 1952, A. R. Brooks, L. A. Konotopetz, including, C. auratus paratypes, 103, 4♀ (CNC). Grande Prairie, July 26, 1961, A. R. Brooks, including, C. auratus paratypes, 2♀ (CNC). High Prairie, July 16, 1961, A. R. Brooks, including C. auratus paratypes, 83, 10♀ (CNC). Hines Creek, June 16, 1961, A. R. Brooks, C. auratus paratypes, 2♀ (CNC). Kananaskis Highway, July 25, 1973, L. A. Kelton, 3♀ (CNC). Kananaskis Road, July 20, 1974, L. A. Kelton, 13 (CNC). Lac la Biche, June 25, 1961, A. R. Brooks, C. auratus paratype, 1♀ (CNC). McMurray, June 30, 1973, W. J. Brown, including C. auratus paratypes, 13, 2♀ (CNC). Nordegg, July 31, 1921, J. McDunnough, C. auratus paratype, 13 (CNC). Robb, August 30, 1970, L. A. Kelton, 1♀ (CNC). Rycroft, August 7, 1961, A. R. Brooks, C. auratus paratype, 1♀ (CNC). Valleyview, August 10, 1961, A. R. Brooks, C. auratus paratype, 13, 1♀ (CNC). British Columbia : Alaska Highway, km 1204, July 30, 1982, G. G. E. Scudder, 13, 1♀ (UBC). Cache Creek, 450–500 m, June 1, 1978, N. L. H. Krauss, 1♀ (AMNH). Chetwynd, 30 km SW of, June 22, 1982, E. Bijdemast, S. G. Cannings, 13 (UBC). Chetwynd, 85 km SW of, on Rt 97, August 26, 1993, M. D. Schwartz, Anaphalis margaritacea (Asteraceae) , 13 (CNC). 3 km S of Chilkat Pass Summit, July 6, 1983, G. G. E. Scud­ der, 13, 4♀ (UBC). Fernie, July 23, 1959, L. A. Kelton, C. auratus paratypes, 53, 9♀ (CNC). Ladysmith, August 14–17, 1970, L. A. Kelton, 13 (CNC). Monaskie Pass, July 16, 1975, L. A. Kelton, 13 (CNC). Pink Mountain, August 17, 1982, L. A. Kelton, 63, 2♀ (CNC). Prophet River Provincial Park, June 24, 1985, E. Bijdemast, 13, 1♀ (UBC). Sicamous, July 17, 1970, L. A. Kelton, 2♀ (CNC). Summit Lake, mi 392 on Alaska Highway, 4500 ft, July 8, 1959, R. E. Leech, 13 (CNC); Summit Lake, mi 392 on Alaska Highway, 4200 ft, July 19–21, 1959, R. E. Leech, 1♀ (CNC). Taylor Flats, July 26, 1948, H. Mac­ Mahon, 33, 4♀ (CNC). Terrace, July 9, 10, 13, 15, 17, 1960, W. R. Richards, 83, 10♀ (CNC). Vernon, June 28, July 2, 1927, D. B. Waddell, 23 (CNC). Manitoba: Clear Lake, July 1, 1930, C. T. Parsons, 13 (AMNH). Falcon Lake, June 1, 25, 1972, L. A. Kelton, 33, 5♀ (CNC). Horton, July 25, 1953, Brooks and Kelton, 1♀ (CAS). Horton, July 25, 1953, Brooks, Wallis, including C. auratus paratypes, 63, 1♀ (CNC). Keld, August 8, 1937, R. H. Beamer, 1♀ (KU). Moose Lake, July 25, 1972, L. A. Kelton, 43, 4♀ (CNC). Riding Mountain National Park, July 20, 1972, L. A. Kelton, 23, 3♀ (CNC). Roblin, 10 mi W of, July 15, 1954, Brooks, Wallis, 23, 1♀ (CNC). Roblin, 30 mi N of, July 13, 14, 1954, Brooks, Wallis, Lathyrus sp. (Fabaceae) , including C. auratus paratypes, 133, 8♀ (CNC). Turtle Mountain, July 21, 1953, Brooks, Wallis, including C. auratus paratype, 13, 1♀ (CNC). New Brunswick: Andover, June 22, 1966, L. A. Kelton, 63, 3♀ (CNC). Chatham, June 27, 1966, L. A. Kelton, Vicia sp. (Fabaceae) , 143, 2♀ (CNC). Edmundston, June 29, July 1–2, 1966, L. A. Kelton, Vicia sp. (Fabaceae) , 163, 17♀ (CNC). Glencoe, June 29, 1966, L. A. Kelton, 13, 2♀ (CNC). Kouchibouguac National Park, June 23, 1978, July 25, 1977, L. A. Kelton, D. J. Brown, J. R. Vockeroth, 83, 4♀ (CNC). Petersville, July 5, 1966, L. A. Kelton, Vicia sp. (Fabaceae) , 143, 11♀ (CNC). White’s Brook, June 29, 1966, L. A. Kelton, 23, 1♀ (CNC). Wolf Lake, August 13, 1966, L. A. Kelton, 1♀ (CNC). Woodstock, June 22, 1966, L. A. Kelton, 1♀ (CNC). Newfoundland: Badger, July 31, 1980, L. A. Kelton, 103, 5♀ (CNC). Change Island, August 1, 1981, D. J. Larson, 1♀ (CNC). Norris Point, Bonne Bay, August 10, 1982, D. J. Larson, 33, 1♀ (CNC). Nova Terra National Park, July 17, 1980, L. A. Kelton, 33 (CNC). Saint John’s, July, 23, 1949, W. J. Brown, 13 (CNC). Northwest Territories: Aklavik, July 16, 1931, Bryant, 1♀ (CAS). Inuvik, July 25, 1983, L. A. Kelton, Artemisia sp. (Asteraceae) , 13, 5♀ (CNC). Nova Scotia: Bible Hill, July 12, 1966, L. A. Kelton, 83, 7♀ (CNC). Cape Breton National Park, August 22, 1966, L. A. Kelton, 153, 18♀ (CNC). Kentville, July 10, 1976, L. A. Kelton, 13 (CNC). Mount Uniacke, July 13, 1966, L. A. Kelton, 93, 4♀ (CNC). Ontario: Algonquin Provincial Park, June 18, 1922, J. Mc­ Dunnough, 13 (CNC). Black Rapids, Ottawa, June 22, 1913, W. Metcalfe, 1♀ (CNC). Black Sturgeon Lake, August 1, 1956, Lindberg, 163, 12♀ (CNC). Collingwood, June 13, 1962, Kelton, Thorpe, C. auratus paratype, 13 (CNC). Devlin, 6 mi S of, June 28, 1960, Kelton, Whitney, 33, 1♀ (CNC). Dryden, August 12, 1960, Kelton, Whitney, 1♀ (CNC). Eagle River, August 11, 1960, Kelton, Whitney, C. auratus paratypes, 33, 1♀ (CNC). Elmvale, June 13, 1962, Kelton, Thorpe, 103, 3♀ (CNC). Emo, July 31, 1960, Kelton, Whitney, C. auratus paratype, 13 (CNC). Emo, 10 mi N of, June 28, 1960, Kelton, Whitney, 113, 3♀ (CNC). Finland, June 28, 1950, Kelton, Whitney, including C. auratus paratypes, 53, 3♀ (CNC). Flower Station, June 28, 1987, D. J. E. Brown, 13, 1♀ (CNC). Fort Frances, July 30–31, 1960, Kelton, Whitney, C. auratus paratypes, 23 (CNC). Fort Frances, July 20, 1969, Kelton and Whitney, 13 (CNC). Grenadier Island Centre, Saint Lawrence Islands National Park, July 27, 1975, E. Wilson, 13 (CNC). Harris Hill, June 24, 1960, Kelton, Whitney, 33 (CNC). Iroquois Falls, June 5, 1987, J. R. Vockeroth, ground among grass roots, 1♀ (CNC). Kapuskasing, July 18, 1959, G. Brumpton, including C. auratus paratypes, 73, 3♀ (CNC). Kenora, 14 mi E of, August 9, 1960, Kelton, Whitney, including C. auratus paratype, 13, 1♀ (CNC). Kirkland Lake, July 17, 1961, G. Brumpton, including C. auratus paratypes, 23, 2♀ (CNC). Long Swamp, Marmora, July 21, 1952, B. P. Beirne, 1♀ (CNC). Marmora, June 25, 1952, J. R. Vockeroth, C. auratus paratype, 1♀ (CNC). Miller Lake, June 12, 1962, Kelton, Thorpe, C. auratus paratypes, 63 (CNC). North Bay, June 19, 1963, L. A. Kelton, including C. auratus paratypes, 123, 5♀ (CNC). Norway Point, Lake of Bays, June 28, 1922, J. Mc­ Dunnough, 1♀ (CNC). One Sided Lake, June 27, 29, August 1, 1960, Kelton and Whitney, including C. auratus paratypes, 313, 32♀ (CNC). Ottawa, July 15, 1957, J. E. H. Martin, C. auratus paratype, 1♀ (CNC). Ottawa, July 2, 1912, E. P. Van Duzee, 53, 3♀ (CAS). Prince Edward County, July 25, 1926, Brimley, 1♀ (CNC). Rainy Riv­ er, June 24, 25, 1960, Kelton, Whitney, including C. auratus paratype, 43, 2♀ (CNC). Sault Ste. Marie, July 5–6, 1961, G. Brumpton, including C. auratus paratypes, 33, 3♀ (CNC). Sibley Park, August 1, 1956, Lindberg (CNC). Sioux Narrows, June 25, 1960, Kelton, Whitney, C. auratus paratype, 13 (CNC). South Gloucester, Blossom Park, Greenbelt, June 9, 1991, M. D. Schwartz, Potentilla argentea (Rosaceae) , 93, 1♀ (CNC). Thomasburg, August 24, 1973, D. G. Reid, Solidago sp. (Asteraceae) , 1♀ (CNC). Wawa, July 7, 1961, G. Brumpton, including C. auratus paratypes, 33, 1♀ (CNC). Prince Edward Island: Brackley Beach, Rustico, July 23, 1967, J. E. H. Martin, 13 (CNC). Cavendish National Park, July 9, 1966 and August 13–19, 1976, L. A. Kelton, 143, 11♀ (CNC). Charlottetown, August 7, 1976, L. A. Kelton, 13, 1♀ (CNC). Dalvey, July 10, 1966, L. A. Kelton, 13 (CNC). Woods Island, July 11, 1966, L. A. Kelton, 93, 4♀ (CNC). Quebec: Bradore Bay, July 29, 1930, W. J. Brown, C. auratus paratype, 1♀ (CNC). Cascapedia, August 17, 1933, W. J. Brown, 1♀ (CNC). Duchesnay, July 5, 1953, O. Peck (CNC), 1♀. Fabre, June 13, July 12, 1963, L. A. Kelton, W. Gagne, Hieracium sp. (Asteraceae) , including C. auratus paratypes, 123, 4♀ (CNC). Gatineau National Park: Browns Lake, Camp Fortune, and north end, June, 11, 25, July 4, 1981, D. J. E. Brown, 193, 19♀ (CNC). Graceville, June 25, 1937, O. Peck, 1♀ (CNC). Harrington Lake, Gatineau National Park, June 18, 1982, D. Brown, 33, 1♀ (CNC). Hudson Heights, August 24, 1956, Lindberg, 23, 3♀ (CNC). Hull, July 16, 1955, L. A. Kelton, 13, 1♀ (CNC). Kingsmere, Gatineau National Park, July 25, 1914, C. G. Hewitt, 13 (CNC). Knowlton, July 8, 1929, L. J. Milne, 13 (CNC). La Minerve, August 28, 1956, Lindberg, 1♀ (CNC). Lac a la Tortue, June 28, 1951, E. G. Munroe, 13, 1♀ (CNC). Lac Mondor, Ste. Flore, June 24, 1951, E. G. Munroe (CNC), 1♀. Ladysmith, July 24, 1958, L. A. Kelton, C. auratus paratype, 1♀ (CNC). Laniel, June 12, 13, 18–19, 24–27, 29, July 1–4, 1963, L. A. Kelton, W. Gagne, Aster sp. (Asteraceae) including C. auratus paratypes, 933, 50♀ (CNC). Laniel, June 5, 1963, W. Gagne, 23 (UCD). Mount Albert, July 11, 1954, W. J. Brown, 13 (CNC). Mount Lyall, 1500 ft, July 3, 25, 1933, W. J. Brown, including C. auratus paratypes, 83, 5♀ (CNC). Old Chelsea, June 11, 1959, J. R. Vockeroth, 13 (CNC). Otter Lake, July 24, 1958, L. A. Kelton, C. auratus paratypes, 33, 3♀ (CNC). Quyon, July 22, 1958, E. G. Munroe, 13, 6♀ (CNC). Thurso, August 20, 1958, L. A. Kelton, C. auratus paratype, 1♀ (CNC). Ville Marie, June 25, 1963, L. A. Kelton, C. auratus paratypes, 33, 3♀ (CNC). Saskatchewan: Asquith, July 19, 1951, L. A. Konotopetz, Anemone canadensis (Ranunculaceae) , including C. auratus paratypes, 13, 2♀ (CNC). Big River, July 3, 1959, A. and J. Brooks, including C. auratus paratypes, 103, 16♀ (CNC). Big River, July 3, 1959, A. and J. Brooks, 1♀ (CAS). Christopher Lake, July 11, 1950, A. R. and J. E. Brooks, 33, 3♀ (CNC). Garrick, June 29, 1950, L. A. Konotopetz, Medicago sativa (Fabaceae) , including C. auratus paratypes, 13, 4♀ (CNC). Kenosee, July 7, 19, 1958, A. R. and J. E. Brooks, including C. auratus paratype, 23 (CNC). Lebret, July 5, 1951, L. A. Konotopetz, 13 (CNC). Love, July 5, 1950, L. A. Konotopetz, Medicago sativa (Fabaceae) , 13 (CNC). Somme, July 8, 1954, Brooks, Wallis, 1♀ (CNC). Torch River Campground on Rt 106, July 12, 1990, M. D. Schwartz, 13 (CNC). Torch River, August 14, 1950, L. A. Konotopetz, Lathyrus venosus (Fabaceae) , including C. auratus paratypes, 23, 2♀ (CNC). White Fox , June 3, 4, 6, 8, 28, 1950, L. A. Konotopetz, 243, 7♀ (CNC). Wood Mountain, August 5, 1955, A. R. Brooks, C. auratus paratype, 13 (CNC). Yukon Territory: Arctic Circle, Dempster Highway, July 29, 1983, L. A. Kelton, 103, 8♀ (CNC). Dawson, July 21, 1982, L. A. Kelton, Artemisia frigida (Asteraceae) 13, 2♀ (CNC). Eagle Plains, July 7, 1983, L. A. Kelton, 13 (CNC). Engineer Creek, Dempster Highway, July 27, 1983, L. A. Kelton, 23, 1♀ (CNC). Junction of Dempster and Klondike Highway, July 20, 1982, L. A. Kelton, 83, 6♀ (CNC). Rancheria, August 11, 1982, L. A. Kelton, Lathyrus sp. (Fabaceae) , 1♀ (CNC). Snag Junction, July 22, 1982, L. A. Kelton, 1♀ (CNC). Tombstone, Dempster Highway, July 27, 1983, L. A. Kelton, 13 (CNC). FINLAND: Tvärminne, August 12, 13, 1960, G. G. E. Scudder, 93, 26♀ (CNC). SWEDEN: Abisko, Lapland, July 29, 31, August 19, 20, 1951, J. R. Vockeroth, 203, 20♀ (CNC). Masugnsbyn, July 23, 1951, J. R. Vockeroth, 13 (CNC). Uppsala, August 6, 1960, G. G. E. Scudder, 2♀ (CNC). USA: Alaska: Kenai Peninsula, 6 mi N of Granite Creek Campground, June 29, 1957, R. W. Fredrickson, 13 (KU). Bristol Bay Co.: Naknek, July 21, 1952, W. R. Mason, 1♀ (CNC). Kenai Peninsula Co.: Moose Pass, 1500 ft, July 2, 1951, W. J. Brown, 13 (CNC). Nome Co.: Unalakleet, August 8, 10, 1961, R. Madge, 23, 6♀ (CNC). South Fairbanks Co.: Chicken, July 22, 1982, L. A. Kelton, 23 (CNC). Tok, July 22, 1982, L. A. Kelton, 13 (CNC). Arizona: Apache Co. : Greer, August 1, 1941, L. L. Stitt, C. arcuatus paratypes, 23, 1♀ (CNC). Greer, August 1, 1941, L. L. Stitt, 1♀ (USNM). Greenlee Co.: Hannagan, Apache National Forest, 9000 ft, July 10, 1968, L. A. Kelton, 103, 8♀ (CNC). California: Nevada Co. : 5 mi N of Hobart Mills, Sagehen Creek Research Station, June 19, 1983, C. B. Barr, 1♀ (LSU). Sagehen Creek, July 5, 1978, M. Williams, 13 (UCD). Siskiyou Co.: Mount Shasta, Shasta National Forest, McBride Springs Campground, June 23, 1985, C. B. Barr, 23, 3♀ (LSU). Colorado: Archuleta Co.: Pagosa Spring, July 22, 1968, L. A. Kelton, 13 (CNC). Clear Creek Co.: Chicago Creek, 8800 ft, August 5, 1961, B. H. Poole, 1♀ (CNC). Doolittle Ranch, Mount Evans, 9800 ft, July 22, 1961 and August 10, 1961, B. H. Poole, J. R. Stainer, 73, 2♀ (CNC). Conejos Co.: Stunner Pass near Platorb, July 27, 1972, H. R. Burke, 1♀ (TAMU). Dolores Co.: Lizard Head, August 1, 1900, E. D. Ball, 1♀ (UAZ). Eagle Co.: 22 km N of Wolcott, Big Piney Creek, June 27, 1989, D. A. Polhemus, Artemisia tridentata (Asteraceae) , 1♀ (JTP). N of Minturn, Grouse Creek Trail, 7840 ft, June 24, 1987, D. A. Polhemus, 113, 1♀ (JTP). Vail, June 23, 1986, J. T. Polhemus, 13 (JTP). Gilpin Co.: Jumbo, Grand Mesa National Forest, 9800 ft, August 25, 1968, L. A. Kelton, 83, 7♀ (CNC). Gunnison Co.: 2 mi N of Gothic, 9300 ft, August 13, 1987, T. J. Henry, 53, 15♀ (USNM). Jackson Co.: 2 mi E of Gould, August 5, 1975, J. C. Schaffner, 1♀ (TAMU). Gould, 9200 ft, August 12, 1968, L. A. Kelton, Artemisia sp. (Asteraceae) , 2♀ (CNC). Muddy Pass, Routt National Forest, 8600 ft, August 19, 1968, L. A. Kelton, Artemisia sp. (Asteraceae) , 4♀ (CNC). Rabbit Ears Pass, August 3, 1947, R. H. Beamer, 13, 3♀ (KU). Park Co.: Geneva Park, 9600 ft, September 6, 1982, D. A. and J. T. Polhemus, 1♀ (JTP). Pitkin Co.: Aspen, 8000 ft, July 24, 1919, 1♀ (AMNH). Routt Co.: Meadows, Routt National Forest, August 20, 1968, L. A. Kelton, Ranunculus sp. (Ranunculaceae) , 103, 15♀ (CNC). Steamboat Springs, 7000 ft, July 16, 1964, H. H. Knight, C. arcuatus paratype, 1♀ (CNC). Steamboat Springs, August 22, 1968, L. A. Kelton, 6♀ (CNC). Teller Co.: 5 mi S of Divide, 9500 ft, August 1, 1955, Ross and Ross, 1♀ (CNC). Connecticut: Litchfield Co.: 4.5 mi N of Salisbury, Mount Riga State Park, July 25, 1970, J. Slater and J. Harrington, 1♀ (AMNH). Idaho: Blaine Co.: Ketchum, July 30, 1955, B. P. Bliven, 43, 4♀ (CAS). Cassia Co. : Burley, July 6, 1931, M. W. Sanderson, 1♀ (KU). Custer Co.: 14 mi NW of Stanley, Vader Creek Camp, 6750 ft, July 14, 1973, C. Musgrave, 13 (OSU). Latah Co.: 5 mi E of Harvard, Palouse River, July 3, 1966, W. Gagne, J. Haddock, 83, 7♀ (CNC). Maine: Kent Co.: E Parsonfield, June 17, 1986, Larochelle, Larivière, 1♀ (CNC). New Hampshire: Coos Co.: Bretton Woods, June 20, 1909, E. P. Van Duzee, 13 (CAS). Mount Washington, Subalpine, July 2, 1896, F. Blanchard, 13 (AMNH). New Mexico: Lincoln Co.: Apache, Sierra Blanca Ski area, 9600 ft, June 20, 1998, E. Riley, 13, 4♀ (TAMU). Otero Co.: 3 mi E of Cloudcroft, July 19, 1976, J. D. Pinto, Geranium sp. (Geraniaceae) , 2♀ (UCR). 4 mi E of Cloudcroft, August 18, 1979, Delorme, McHugh, Schaffner, 33, 8♀ (TAMU). Cloudcroft, July 23, 1967, L. A. Kelton, 3♀ (CNC). Cloudcroft, June 27, 1940, D. E. Hardy and R. H. Beamer, 73, 9♀ (KU). New York: Cattaraugus Co.: Rock City, July 5, 1915, H. H. Knight, C. auratus paratypes, 43, 1♀ (AMNH, CNC). Essex Co.: Whiteface Mountain, 4600– 4872 ft, July 19, 1962, J. R. Vockeroth, 13 (CNC). Pennsylvania: Centre Co.: Scotia Barrens, June 25, 1988, A. G. Wheeler, Jr., 13, 17♀ (PDA). Scotia Barrens, May 19, 1980, A. G. Wheeler, Jr., Stellaria longifolia (Caryophyllaceae) , 123, 15♀ (PDA, AMNH). Utah: Duchesne Co.: Indian Canyon, July 14, 1964, G. F. Knowlton, 13 (UCD). Sevier Co.: Fish Lake National Forest, R2E, T25S, sec. 24, 25, 8720 ft, June 16, 1980, G. M. Stonedahl, Artemisia sp. (Asteraceae) , 23, 1♀ (OSU). Vermont: Grand Isle Co.: Grand Isle, July 3, 1900, A. P. Morse, 1♀ (AMNH).

Figures

Fig. 1. Habitus views of Chlamydatus spp.: C. artemisiae–C. opacus. See appendix 1 for localities.

Fig. 3. Coloration patterns on legs of Chlamydatus spp.

Fig. 5. Male genitalia of Chlamydatus spp.: C. monilipes–C. pulicarius. See appendix 1 for localities.

Fig. 2. Habitus views of Chlamydatus spp.: C. pallidicornis–C. wilkinsoni. See appendix 1 for localities.

Fig. 9. Scanning electron micrographs of Chlamydatus pulicarius, 3, British Columbia: Pink Mountain. A. Head, frontal view. B. Head, lateral view. C. Setae on hemelytra. D. Pretarsus, frontal view. E. Mesothoracic spiracle and metaepisternal scent efferent system, lateral view. F. Pygophore, posterolateral view, showing left paramere and phallotheca in repose. Scales as indicated.