Sphaerocionium

Ahamada H. Saïd, Sabine Hennequin, Germinal Rouhan & Jean-Yves Dubuisson, 2017, Disentangling the diversity and taxonomy of Hymenophyllaceae (Hymenophyllales, Polypodiidae) in the Comoros, European Journal of Taxonomy 313, pp. 1-53 : 22

Subgenus Sphaerocionium

Subgenus Sphaerocionium is clearly distinct in the archipelago with two species characterized by lamina covered by a dense indumentum (see Fig. 2A ). By contrast, all other Comorian Hymenophyllum species display glabrous lamina margins. Furthermore, the two Comorian Sphaerocionium species, Hymenophyllum hirsutum (L.) Sw. and H. capillare are easily distinguished from each other: H. capillare usually has wingless stipes and rachis (except in dwarf forms exhibiting winged rachis, at least in their upper part) while H. hirsutum always bears a developed wing all along the stipes and rachis. In addition, lamina hairs are pedunculated in H. hirsutum (see Fig. 2A ) and not pedunculated in H. capillare .

Figures

Fig. 2. Illustration of representative descriptors and states used in the Xper database (see Appendix 1). A. Hymenophyllum hirsutum (L.) Sw., typical bilabiate sori (S) without a developed base, fully hairy lamina with pedunculated brown trichomes (see inset). B. Didymoglossum robinsonii (Hook. ex Baker) Copel., false veins (FV) parallel but not connected to true veins (TV) and marginal dark trichomes clustered here by 6 (see inset). C. Crepidomanes inopinatum (Pic. Serm.) J. P. Roux, typical trichomanoid sori (S) with a developed tubular base, lips are here dilated as a collar. D. Didymoglossum rotundifolium (Bonap.) J. P. Roux, false veins (FV) parallel but not connected to true veins (TV) and continuous submarginal false vein (smFV), here the one-cell thick lamina appears well translucent. E. Crepidomanes bipunctatum (Poir.) Copel., false veins more or less continuous and parallel to margins (mFV) and not connected to true veins (TV). F. Hymenophyllum sibthorpioides (Bory ex Willd.) Mett. ex Kuhn, bilabiate sori (S) with toothed margins. Photographs by A. H. Saïd.