Cybaeus ubicki Bennett,

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade, Zootaxa 4965 (3), pp. 401-436: 424

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Cybaeus ubicki Bennett

spec. nov.

Cybaeus ubicki Bennett  spec. nov.

Figs 68–79View FIGURES 68–71View FIGURES 72–73View FIGURES 74–79, 84View FIGURE 84

Type material. U.S.A.: California: Holotype male, Marin County, S.P. Taylor State Park , 8.xi.1953, V. D. Roth & R. O. Schuster ( CAS)  . Paratypes: Marin. 1♂, Burdell Mtn. , 12.i.1984, D. Ubick ( CAS)  ; 1♀, SE slope of Burdell Mtn. , 16.iii.1990, D. Ubick & T. Briggs ( CAS)  ; 4♀, Inverness , 8.xi.1953, V. D. Roth ( CAS)  ; 1♀, Mt. Tamalpais State Park, near Cataract Falls , 12.ii.2013, I.M. Sokolov ( CAS)  ; 2♂ 2♀, 4 mi. E of Muir Woods National Monument , 10.i.1964, V.D. Roth ( CAS)  ; 3♂ 1♀, Novato , 7.i.1987, D. Ubick ( CAS)  ; 3♂ 12♀, Novato, San Marin Dr. , 18.xii.1982, D. Ubick ( CAS)  ; 3♂ 5♀, Novato, San Marin Dr. , 7.xii.1985, D. Ubick ( CAS)  ; 4♀, Novato, San Marin Dr. , 2.xii.1988, D. Ubick ( CAS)  ; 1♂, Novato , open space N of San Marin Dr., 28.xi1987, D. Ubick ( CAS)  ; 1♂, Novato , open space N of Simmon’s Lane, 4.i.1990, D. Ubick & A. Gudeli ( CAS)  ; 1♀, 7 mi. E of Point Reyes Station , 1.iii.1960, Smith & Schuster ( AMNH)  ; 3♂ 2♀, S.P. Taylor State Park , 8.xi.1953, V. D. Roth & R. O. Schuster ( CAS)  ; 1♀, S end of S.P. Taylor State Park, 1.xi.1953, V. D. Roth & G. Marsh ( CAS)  ; 1♂ 2♀, World College West, 1 mi. W of highway 101, San Antonio Rd. , 60 m elev., 11.i.1986, D. Ubick ( CAS)  ; Mendocino   . 3♂ 2♀, 5 mi. E of Anchor Bay , 12.ix.1961, W.J. Gertsch & W. Ivie ( AMNH)  ; Sonoma   . 1♀, Armstrong Redwoods State Natural Reserve , 14.iii.1954, J. Helfer ( CAS)  ; 3♀, 2.1 mi. NW of Camp Meeker , 90 m elev., 14.i.1983, T. S. Briggs, V. F. Lee, & D. Ubick ( CAS)  ; 1♀, Kruse Rhododendron State Natural Reserve , 9.x.1954, Schuster & MacNeill ( CAS)  ; 1♀, Monte Rio ,, P. Craig, W. Savary, & D. Ubick ( CAS)  ; 2♀, 2.4 mi. SE of Monte Rio , 20.xii.2001, T. Briggs, G. Giribet, D. Ubick, & S. Ubick ( CAS)  ; 1♀, Salt Point State Park Campground, 29.iii.1980, D. Ubick ( CAS)  .

Etymology. The specific epithet is a patronym honoring Darrell Ubick whose unconditional help and prodigious collecting expertise has greatly benefited our work on Californian Cybaeidae  ; name in the genitive case.

Diagnosis. The male of C. ubicki  spec. nov. is unlikely to be confused with the males of the other species in the consocius  group and is distinguished by a combination of features of the patellar apophysis and the genital bulb. The patellar apophysis ( Figs 68–69View FIGURES 68–71, 72View FIGURES 72–73) is relatively short (length about 1/3 width of the patella) with the peg setae divided into basal and distal groups; the basal group is clustered on a small dorsal prominence (in some specimens the separation into two groups is indistinct). The embolus is broad and flattened medially ( Figs 70–71View FIGURES 68–71) and the proximal arm of the tegular apophysis ( Figs 70–71View FIGURES 68–71, 73View FIGURES 72–73) features a deep ventral longitudinal groove and terminates in a broad rounded tip. In addition the palpal tibia bears a small flattened ridge (inconspicuous in some specimens) dorsal to the retrolateral tibial apophysis ( Figs 68–69View FIGURES 68–71). Among the females of the consocius  group, the female of C. ubicki  spec. nov. is most likely to be confused with the other species which lack U-shaped copulatory ducts: C. opulentus  spec. nov., C. penedentatus  , C. vulpinus  , C. pan  spec. nov., and C. simplex  . The female of C. ubicki  spec. nov. is characterized by its large transverse atrium (nearly as wide as the vulva) ( Figs 74–75, 77–78View FIGURES 74–79) and the broad, conspicuously prominent copulatory ducts ( Figs 75–76, 78–79View FIGURES 74–79). Separating the female of C. ubicki  spec. nov. from the females of C. opulentus  spec. nov., C. pan  spec. nov., and C. simplex  is discussed under the diagnoses of those species. The female of C. vulpinus  has a large atrium but it is only about 1/2 the width of the vulva ( Fig. 40View FIGURES 37–41) and the copulatory ducts are narrower and relatively inconspicuous ( Fig. 40–41View FIGURES 37–41). The female of C. penedentatus  has a transverse atrium but it is very small ( Figs 46–47, 49–50View FIGURES 46–51, 52View FIGURES 52–54).

Description. Ventral tibia I macrosetae usually 2-1p-2-1p-0.

Male: (n=22). Patellar apophysis ( Figs 68–69View FIGURES 68–71, 72View FIGURES 72–73) with about 12–15 peg setae usually in two groups; groups merge in some specimens). Proximal arm of tegular apophysis ( Figs 70View FIGURES 68–71, 73View FIGURES 72–73) with small excavation on posterior margin near tip. Measurements (n=19). CL 1.80–2.8 (2.2+0.3), CW 1.35–1.98 (1.62+0.20), SL 0.91–1.30 (1.10+0.13), SW 0.91–1.21 (1.05+0.11). Holotype CL 2.13, CW 1.50, SL 1.04, SW 1.00.

Female: (n=47). Atrium ( Figs 74–75, 77-78View FIGURES 74–79) medially located on epigynum. Copulatory ducts ( Figs 75–76, 78– 79View FIGURES 74–79) converging along midline anterior to atrium then diverging anteriorly before turning posteriorly. Spermathecal stalks contiguous ( Fig. 76View FIGURES 74–79) or widely separated ( Fig. 79View FIGURES 74–79) near Bennett’s glands. Measurements (n=22). CL 1.73–2.6 (2.2+0.3), CW 1.25–1.73 (1.52+0.17), SL 0.90–1.26 (1.11+0.12), SW 0.83–1.18 (1.03+0.11).

Distribution and natural history. Coastal central California from southern Mendocino County to southern Marin County ( Fig. 84View FIGURE 84). Males have been collected from November through January and in September.


Royal British Columbia Museum - Herbarium


Departamento de Geologia, Universidad de Chile


California Academy of Sciences


Tavera, Department of Geology and Geophysics


American Museum of Natural History