Cybaeus simplex Roth,

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade, Zootaxa 4965 (3), pp. 401-436: 420-423

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Cybaeus simplex Roth


Cybaeus simplex Roth 

Figs 1View FIGURES 1–4, 59–67View FIGURES 59–63View FIGURES 64–67, 85View FIGURE 85

Cybaeus simplex Roth, 1952: 216  , figs 9–12. Roewer 1954: 92. Roth and Brown 1986: 4. World Spider Catalog 2021

Type material examined. U.S.A.: Oregon: Holotype male, Linn County, Cascadia , 11.ix.1948, V. D. Roth ( AMNH), subsequently lost  . Neotype male here designated from original paratype series, Linn County, Cascadia , 5.ix.1948, V. D. Roth ( AMNH)  . Neotype lacks its left palpus. Allotype female, Linn County, Cascadia , 11.ix.1948, V. D. Roth ( AMNH), subsequently lost  .

Other material examined. U.S.A.: Oregon: Coos. 2♀, 3 mi. N of Bandon , 30.ix.1959, V. D. Roth ( CAS);  Jackson. 1♂, Ashland Watershed , 11.x.1998, R. W. Peck ( RBCM);  Josephine. 1♂, Oregon Cave, Oregon Cave National Monument, 24.x.2011, N. Mason ( NHM);  Lane. 1♀, Blue Pool Camp , Willamette National Forest , 7.ix.1949, V.D. Roth ( AMNH);  2♂, H.J. Andrews Experimental Forest, 18.x–9.xi.1983, J.D. McIver ( CNC);  Linn. 1♂ 1♀, Cascadia , 5.ix.1948, V.D. Roth ( AMNH);  1♂ 1♀, Cascadia , 5.ix.1948, V.D. Roth ( MCZ);  1♂ 1♀, Cascadia , 11.ix.1948, V.D. Roth ( NMNH);  1♀, 1 mi. E of Quartzville (27 mi. NE Sweet Home ), 7.iii.1948, V.D. Roth ( CAS)  .

Diagnosis. The male of C. simplex  is unlikely to be confused with the males of the other species in the consocius  group except for C. opulentus  . Distinguishing the males of these two species is discussed under the diagnosis of C. opulentus  . Among the females of the consocius  group, the female of C. simplex  is most likely to be confused with the other species which lack U-shaped copulatory ducts: C. ubicki  spec. nov., C. penedentatus  , C. vulpinus  , C. pan  spec. nov., and C. opulentus  spec. nov. From those species the female of C. simplex  is distinguished by a combination of its wide and arched transverse atrium, about 2/3 the width of the vulva ( Figs 64–65View FIGURES 64–67) (versus atrium much smaller, about 1/3 the width of the vulva in C. penedentatus  [ Figs 46–47View FIGURES 46–51, 52–53View FIGURES 52–54]); the relatively narrow lumina of its copulatory ducts ( Figs 65–67View FIGURES 64–67) (versus broad copulatory duct lumina, especially at the junction of the copulatory ducts with the atrium, in C. ubicki  spec. nov. [ Figs 75–76View FIGURES 74–79], C. opulentus  spec. nov. [ Fig. 31View FIGURES 30–32], C. pan  spec. nov. [ Fig. 38View FIGURES 37–41], and C. vulpinus  [ Fig. 40View FIGURES 37–41]); copulatory ducts contiguous at the atrium as well as partway between the atrium and the spermathecal heads ( Figs 65–67View FIGURES 64–67) (versus contiguous only at the atrium in C. pan  spec. nov. [ Fig. 38View FIGURES 37–41] and C. vulpinus  [ Fig. 40View FIGURES 37–41]); and the relatively long copulatory ducts and spermathecal stalks ( Figs 65–67View FIGURES 64–67) (versus short copulatory ducts and spermathecal heads in C. opulentus  spec. nov. [ Fig. 32View FIGURES 30–32]).

Description. Roth (1952) described both sexes in detail. Ventral tibia I macrosetae: 2-1p-2-1p-2.

Male: (n=9). Patellar apophysis ( Figs 1View FIGURES 1–4, 60View FIGURES 59–63) about as long as width of patella with peg setae clustered around tip and linearly along dorsal surface. Distal extension of retrolateral tibial apophysis ( Fig. 59View FIGURES 59–63) short, digitiform. Measurements (n=6). CL 2.28–2.6 (2.5±0.1), CW 1.48–1.93 (1.77±0.20), SL 1.17–1.31 (1.26±0.05), SW 1.00–1.21 (1.15±0.08). Holotype, neotype: CL 2.6, 2.6, CW 1.90, 1.93, SL 1.30, 1.30, SW 1.21, 1.20.

Female: (n=8). Atrium ( Fig. 64View FIGURES 64–67) anteriorly located on epigynum. Copulatory ducts ( Figs 65–67View FIGURES 64–67) narrowly contiguous at atrium, projecting anteriorly for short distance from atrium then turning posteriorly and becoming contiguous again along midline before joining spermathecal heads at lateral margins of vulva. Stalks ( Figs 65–67View FIGURES 64–67) often contiguous or nearly so in region of Bennett’s glands. Measurements (n=6). CL 2.15–2.7 (2.4±0.2), CW 1.48–1.78 (1.60±0.13), SL 1.04–1.30 (1.17±0.09), SW 0.92–1.21 (1.07±0.10).

Distribution and natural history. Coos, Jackson, Josephine, Lane, and Linn Counties of western Oregon from the Cascade Range east of the Willamette Valley, west to the coast and south nearly to California ( Fig. 85View FIGURE 85). Mature males have been collected from September to November. The holotype and allotype pair was found “under bark about 1 inch apart.”


Royal British Columbia Museum - Herbarium


American Museum of Natural History


California Academy of Sciences


Departamento de Geologia, Universidad de Chile


Royal British Columbia Museum


University of Nottingham


Canadian National Collection of Insects, Arachnids, and Nematodes


Museum of Comparative Zoology


Smithsonian Institution, National Museum of Natural History














Cybaeus simplex Roth

Bennett, Robb, Copley, Claudia & Copley, Darren 2021

Cybaeus simplex

Roth, V. D. & Brown, W. L. 1986: 4
Roewer, C. F. 1954: 92
Roth, V. D. 1952: 216