Cybaeus pan Bennett,

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade, Zootaxa 4965 (3), pp. 401-436: 414-417

publication ID

publication LSID


persistent identifier

treatment provided by


scientific name

Cybaeus pan Bennett

spec. nov.

Cybaeus pan Bennett  spec. nov.

Figs 33–39View FIGURES 33–36View FIGURES 37–41, 58View FIGURES 55–58, 86View FIGURE 86

Type material. U.S.A.: California: Holotype male, Sierra County, 2 mi. N of Calpine , 6.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)  . Paratypes: Butte. 1♀, Lumpkin Rd. near Feather Falls, Oroville , 18.iv.1994, J. Boutin ( CAS)  ; Plumas   . 1♀, NE of Bucks Lake , 4.ix.1988, D. Ubick ( CAS)  ; Nevada. 1♀, 1 mi. N of Washington , 3050’, 27.xii.1986, D. Ubick ( CAS)  ; Sierra   . 9♂ 10♀, 2 mi N of Calpine , 6.ix.1959, V. D. Roth & W.J Gertsch ( AMNH)  ; 1♀, 5 mi. E of Camptonville , 14.vii.2002, D. Ubick ( CAS)  ; 1♂ 1♀, Sierra City, The Cups , 6.ix.1959, V. D. Roth & W.J Gertsch ( AMNH)  ; 1♀, 1.5 mi. E of Sierra City, Wild Plum Campground , 4200’, 12–16.viii.1996, D. Ubick ( CAS)  ; 1♀, 1.5 mi. E of Sierra City, Wild Plum Campground , 4200’, 9.viii.2001, D. Ubick ( CAS)  ; 1♂, 1.5 mi. E of Sierra City, Wild Plum Campground , 4200’, 18.vii.2002, D. Ubick ( CAS)  ; 2♂, 3.5 mi. NE of Sierra City , 5100’, 17.viii.1993 ’, D. Ubick ( CAS)  ; 2♀, 6 mi. NE of Sierra City, SFSU Field Station , 15–18.viii.1991, D. Ubick ( CAS)  ; 1♂ 5♀, 6 mi. NE of Sierra City, SFSU Field Station , 5500’, 10–18.viii.1992, D. Ubick ( CAS)  ; 1♂ 1♀, 6 mi. NE of Sierra City, SFSU Field Station , 5500’, 16–20.viii.1993, D. Ubick ( CAS)  ; 1♂ 2♀, 6 mi. NE of Sierra City, SFSU Field Station , 5500’, 7–11.viii.2000, D. Ubick ( CAS)  ; 1♀, Yuba Pass , 8.viii.2001, D. Ubick ( CAS)  ; 1♀, Yuba Pass , 13.viii.1997, G. deNevers & H. Wilcox ( CAS)  ; 3♂ 3♀, Yuba Pass , 9.viii.2000, D. Ubick ( CAS)  ; 1♀, Yuba Pass , 8.viii.2001, R. Love ( CAS)  ; 2♀, Yuba Pass ,, D. Ubick ( CAS)  .

Etymology. The specific epithet is a noun in apposition from Pan, the ancient pipe-playing Greek god of the countryside. Cybaeus pan  spec. nov. is named in honor of the late John “Pan” Logan, dear friend and native of the northern Sierra Nevada region of California, who did much in his lifetime to uphold the traditions associated with mythical Pan; name in apposition.

Diagnosis. Among the males of the consocius  group species, C. pan  spec. nov. is only likely to be confused with C. vulpinus  . The males of the two species are distinguished by, in C. pan  spec. nov., a relatively simple, smoothly curved and untwisted proximal arm of the tegular apophysis in ventral view ( Figs 35–36View FIGURES 33–36) with, in prolateral view ( Fig. 58View FIGURES 55–58), a small triangular dorsal keel (versus smoothly curved but twisted proximal arm lacking a dorsal process [ Figs 57View FIGURES 55–58, 82View FIGURES 80–83] in C. vulpinus  ). As well the patellar apophysis ( Figs 33–34View FIGURES 33–36) is longer (about 3/4 the width of the patella), anterolaterally directed, and has more peg setae (about 35) in C. pan  spec. nov. (versus about 1/3 the width of the patella, anteriorly directed, and bears only 13 peg setae in the single known male of C. vulpinus  [ Fig. 81View FIGURES 80–83]). Among the females of the consocius  group, the female of C. pan  is most likely to be confused with the other species which lack U-shaped copulatory ducts: C. ubicki  spec. nov., C. penedentatus  , C. vulpinus  , C. simplex  , and C. opulentus  spec. nov. Among these species, the female of C. pan  spec. nov. is most difficult to separate from C. vulpinus  ; the epigyna and vulvae of these two species are very similar and, in some cases, distribution may be the best or only way to separate them ( Fig. 86View FIGURE 86). The only apparent morphological character separating the two species is the conformation of the spermathecal stalks: in C. pan  spec. nov. they are relatively long, distinctly sinuous and describing an acute angle (approximately 55 ⁰ –65 ⁰) near the Bennett’s glands ( Figs 38–39View FIGURES 37–41) versus in C. vulpinus  ( Figs 40–41View FIGURES 37–41) only slightly sinuous and obtusely angled (circa 120 ⁰ –130 ⁰). Separating the females of C. pan  spec. nov. and C. opulentus  spec. nov. is discussed under the diagnosis of the latter. From C. ubicki  spec. nov. and C. penedentatus  , the female of C. pan  spec. nov. is distinguished by a combination of a large, conspicuous atrium occupying about 1/2 the width of the vulva ( Figs 37–38View FIGURES 37–41) (versus atrium small, inconspicuous, occupying about 1/3 the width of the vulva in C. penedentatus  [ Figs 46–47, 49View FIGURES 46–51, 52View FIGURES 52–54]) and copulatory ducts relatively small, narrow, and inconspicuous ( Figs 38–39View FIGURES 37–41) (versus large, broad, and prominent in C. ubicki  spec. nov. [ Figs 75–76, 78–79View FIGURES 74–79]). Finally, specimens of both sexes of C pan  spec. nov. (as well as of C. vulpinus  ) usually have banded femora; specimens of all the other consocius  group species usually have unbanded femora.

Description. Femora usually banded. Ventral tibia I macrosetae usually 2-1p(or 0)-2-1p-1p(or 0).

Male: (n=20). As per Diagnosis. Measurements (n=9). CL 2.33–2.8 (2.6±0.2), CW 1.70–2.05 (1.90±0.13), SL 1.20–1.38 (1.30±0.06), SW 1.16–1.33 (1.25±0.05). Holotype CL 2.43, CW 1.80, SL 1.22, SW 1.20.

Female: (n=35). Epigynum usually with small notch posteriorly ( Fig. 38View FIGURES 37–41) (inconspicuous and may be difficult to discern, most easily observed in cleared specimens), absent in some specimens ( Fig. 37View FIGURES 37–41); atrium ( Figs 37–38View FIGURES 37–41) medially located on epigynum, usually strongly arched (some specimens have a weakly arched atrium approximating the characteristic atrial morphology seen in specimens of C. vulpinus  [e.g., Fig. 40View FIGURES 37–41]). Copulatory ducts ( Figs 38–39View FIGURES 37–41) separated except at atrium; anteriorly directed and diverging from atrium, anterior margin often indistinct. Stalks of spermathecae simple ( Figs 38–39View FIGURES 37–41). Measurements (n=13). CL 2.20–3.0 (2.5±0.2), CW 1.50–2.05 (1.74±0.16), SL 1.13–1.40 (1.25±0.08), SW 1.05–1.33 (1.19±0.07).

Distribution and natural history. Northern Sierra Nevada region of eastern California northwest of Lake Tahoe in Butte, Plumas, Nevada, and Sierra Counties ( Fig. 86View FIGURE 86). Mature males have been collected from July through September.


Royal British Columbia Museum - Herbarium


American Museum of Natural History


California Academy of Sciences


Departamento de Geologia, Universidad de Chile