Cybaeus opulentus Bennett

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade, Zootaxa 4965 (3), pp. 401-436: 412-414

publication ID

https://doi.org/10.11646/zootaxa.4965.3.1

publication LSID

lsid:zoobank.org:pub:94FB89CF-2083-4FAC-AE60-B8CCF1D5FE8E

DOI

http://doi.org/10.5281/zenodo.4727176

persistent identifier

http://treatment.plazi.org/id/038B87C3-1908-BE22-FF3A-6ECFFDFE997F

treatment provided by

Plazi

scientific name

Cybaeus opulentus Bennett
status

spec. nov.

Cybaeus opulentus Bennett   spec. nov.

Figs 27–32 View FIGURES 27–29 View FIGURES 30–32 , 55 View FIGURES 55–58 , 86 View FIGURE 86

Type material. U.S.A.: California: Holotype male, El Dorado County, 4 mi. W of Kyburz , 15.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)   . Paratypes: Alpine. 1♀, North Fork Mokelumne River at Hwy #4, 7000’, 17.viii.1990, D. Ubick ( CAS)   ; 1♂, Woods Lake , 8300’, 16.viii.1990, D. Ubick ( CAS)   ; El Dorado. 1♂, Bridal Veil Falls , 10.5 mi. E of Camino, 3400’, 4.ix.1999 (matured), D. Ubick & T. Briggs ( CAS)   ; 3♂, Echo Summit , 7377’, 19.ix.1963, W.J. Gertsch ( AMNH)   ; 4♂ 3♀, Echo Summit , 7382’, 9.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)   ; 1♂, 2.ix.1961, W.J. Gertsch & W. Ivie ( AMNH)   ; 2♂ 3♀, Fallen Leaf Lake , 9.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)   ; 2♂ 1♀, 14.4 km. E of Georgetown , 1300 m., 18.ix.2012, C. Pavla & G. Griswold ( CAS)   ; 3♂, 15.2 km. E of Georgetown , 1300 m., 16.ix.2012, C. Pavla & G. Griswold ( CAS)   ; 1♂, Kyburz , 15.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)   ; 1♀, Riverton , 22.ii.1958, R. O. Schuster ( CAS)   ; 2♂ 8♀, 3 mi. W of Riverton , 9.ix.1959, W.J. Gertsch & V.D. Roth ( AMNH)   ; Placer   . 1♀, Baxter , 3900’, 12.vii.1990. D. Ubick ( CAS)   ; 1♂, 6 mi. NE of Foresthill , 3480’, 27.xii.1986, D. Ubick ( CAS)   ; 1♂ 2♀, 9 mi. NE of Foresthill , 3720’, 27.xii.1986, D. Ubick ( CAS)   .

Etymology. The specific epithet is from a Latin adjective meaning rich or wealthy and refers to the distribution of this species in eastern California in the vicinity of Lake Tahoe as well as to the central importance of the area to the California gold rush of the mid nineteenth century.

Diagnosis. The male of C. opulentus   spec. nov. is unlikely to be confused with the males of the other species in the consocius   group and is diagnosed by the unique form of the proximal arm of its tegular apophysis. The base of the proximal arm bears a prominent protrusion and the tip of the proximal arm is slightly twisted ( Figs 28–29 View FIGURES 27–29 , 55 View FIGURES 55–58 ). Among the other males in the consocius   group only C. simplex   is similar. In the male of C. simplex   the proximal arm bears a much more prominent basal protrusion and the tip is strongly twisted ( Figs 1 View FIGURES 1–4 , 61–63 View FIGURES 59–63 ). In addition, the patellar apophysis in the male of C. opulentus   spec. nov. bears more peg setae (30–40 [ Fig. 27 View FIGURES 27–29 ] versus 15–25 in C. simplex   [ Fig. 60 View FIGURES 59–63 ]) and the retrolateral tibial apophysis does not extend distally beyond the end of the tibia ( Fig. 27 View FIGURES 27–29 ) (versus retrolateral tibial apophysis with a significant distal extension in C. simplex   [ Fig. 59 View FIGURES 59–63 ]). Among the females of the consocius   group, the female of C. opulentus   spec. nov. is only likely to be confused with the other species which lack U-shaped copulatory ducts: C. ubicki   spec. nov., C. penedentatus   , C. vulpinus   , C. pan   spec. nov., and C. simplex   . From those species the female of C. opulentus   spec. nov. is distinguished by a combination of atrium wide, occupying about 2/3 the width of the vulva and located on anterior third of the vulva ( Figs 30–31 View FIGURES 30–32 ) (versus atrium much smaller, about 1/3 the width of the vulva, and usually located on the medial third of the vulva in C. penedentatus   [ Figs 46–47 View FIGURES 46–51 , 52–53 View FIGURES 52–54 ] or atrium located on medial third of the vulva and wider, nearly as wide as the vulva, in C. ubicki   spec. nov. [ Figs 74–75, 77–78 View FIGURES 74–79 ]); copulatory ducts and spermathecal stalks short and spermathecal stalks noncontiguous along midline of vulva ( Figs 31–32 View FIGURES 30–32 ) (versus copulatory ducts and spermathecal stalks relatively long and spermathecal stalks contiguous along vulval midline in C. simplex   [ Figs 65–67 View FIGURES 64–67 ]; copulatory ducts contiguous at the atrium as well as partway between the atrium and the spermathecal heads ( Figs 31–32 View FIGURES 30–32 ) (versus contiguous only at the atrium in C. pan   spec. nov. [ Figs 38–39 View FIGURES 37–41 ] and C. vulpinus   [ Figs 40–41 View FIGURES 37–41 ]).

Description. Ventral tibia I macrosetae 2-1p-2-1p-1p(or 2 or 0).

Male: (n=23). Patellar apophysis ( Fig. 27 View FIGURES 27–29 ) relatively short, length about 2/3 width of patella. Tip of proximal arm of tegular apophysis ( Figs 28 View FIGURES 27–29 , 55 View FIGURES 55–58 ) elongate. Measurements (n=15). CL 2.43–3.3 (2.9±0.2), CW 1.80–2.35 (2.09±0.17), SL 1.25–1.58 (1.40±0.09), SW 1.17–1.45 (1.34±0.08). Holotype CL 2.8, CW 1.93, SL 1.33, SW 1.30.

Female: (n=20).Atrium ( Fig. 30 View FIGURES 30–32 ) transverse, arched. Copulatory ducts ( Figs 31–32 View FIGURES 30–32 ) broad at atrium, convergent anteriorly along midline of vulva then turning posteriorly. Measurements (n =18). CL 2.40–3.2 (2.8±0.2), CW 1.60–2.18 (1.94±0.17), SL 1.18–1.50 (1.36±0.09), SW 1.09–1.45 (1.30±0.09).

Distribution and natural history. South central Sierra Nevada region of eastern California south and west of Lake Tahoe in Alpine, El Dorado, and Placer Counties ( Fig. 86 View FIGURE 86 ). Mature males have been collected from August through September and in December.

V

Royal British Columbia Museum - Herbarium

AMNH

American Museum of Natural History

CAS

California Academy of Sciences

T

Tavera, Department of Geology and Geophysics

R

Departamento de Geologia, Universidad de Chile

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Cybaeidae

Genus

Cybaeus