Cybaeus opulentus Bennett,

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade, Zootaxa 4965 (3), pp. 401-436: 412-414

publication ID

publication LSID


persistent identifier

treatment provided by


scientific name

Cybaeus opulentus Bennett

spec. nov.

Cybaeus opulentus Bennett  spec. nov.

Figs 27–32View FIGURES 27–29View FIGURES 30–32, 55View FIGURES 55–58, 86View FIGURE 86

Type material. U.S.A.: California: Holotype male, El Dorado County, 4 mi. W of Kyburz , 15.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)  . Paratypes: Alpine. 1♀, North Fork Mokelumne River at Hwy #4, 7000’, 17.viii.1990, D. Ubick ( CAS)  ; 1♂, Woods Lake , 8300’, 16.viii.1990, D. Ubick ( CAS)  ; El Dorado. 1♂, Bridal Veil Falls , 10.5 mi. E of Camino, 3400’, 4.ix.1999 (matured), D. Ubick & T. Briggs ( CAS)  ; 3♂, Echo Summit , 7377’, 19.ix.1963, W.J. Gertsch ( AMNH)  ; 4♂ 3♀, Echo Summit , 7382’, 9.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)  ; 1♂, 2.ix.1961, W.J. Gertsch & W. Ivie ( AMNH)  ; 2♂ 3♀, Fallen Leaf Lake , 9.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)  ; 2♂ 1♀, 14.4 km. E of Georgetown , 1300 m., 18.ix.2012, C. Pavla & G. Griswold ( CAS)  ; 3♂, 15.2 km. E of Georgetown , 1300 m., 16.ix.2012, C. Pavla & G. Griswold ( CAS)  ; 1♂, Kyburz , 15.ix.1959, W.J. Gertsch & V. D. Roth ( AMNH)  ; 1♀, Riverton , 22.ii.1958, R. O. Schuster ( CAS)  ; 2♂ 8♀, 3 mi. W of Riverton , 9.ix.1959, W.J. Gertsch & V.D. Roth ( AMNH)  ; Placer   . 1♀, Baxter , 3900’, 12.vii.1990. D. Ubick ( CAS)  ; 1♂, 6 mi. NE of Foresthill , 3480’, 27.xii.1986, D. Ubick ( CAS)  ; 1♂ 2♀, 9 mi. NE of Foresthill , 3720’, 27.xii.1986, D. Ubick ( CAS)  .

Etymology. The specific epithet is from a Latin adjective meaning rich or wealthy and refers to the distribution of this species in eastern California in the vicinity of Lake Tahoe as well as to the central importance of the area to the California gold rush of the mid nineteenth century.

Diagnosis. The male of C. opulentus  spec. nov. is unlikely to be confused with the males of the other species in the consocius  group and is diagnosed by the unique form of the proximal arm of its tegular apophysis. The base of the proximal arm bears a prominent protrusion and the tip of the proximal arm is slightly twisted ( Figs 28–29View FIGURES 27–29, 55View FIGURES 55–58). Among the other males in the consocius  group only C. simplex  is similar. In the male of C. simplex  the proximal arm bears a much more prominent basal protrusion and the tip is strongly twisted ( Figs 1View FIGURES 1–4, 61–63View FIGURES 59–63). In addition, the patellar apophysis in the male of C. opulentus  spec. nov. bears more peg setae (30–40 [ Fig. 27View FIGURES 27–29] versus 15–25 in C. simplex  [ Fig. 60View FIGURES 59–63]) and the retrolateral tibial apophysis does not extend distally beyond the end of the tibia ( Fig. 27View FIGURES 27–29) (versus retrolateral tibial apophysis with a significant distal extension in C. simplex  [ Fig. 59View FIGURES 59–63]). Among the females of the consocius  group, the female of C. opulentus  spec. nov. is only likely to be confused with the other species which lack U-shaped copulatory ducts: C. ubicki  spec. nov., C. penedentatus  , C. vulpinus  , C. pan  spec. nov., and C. simplex  . From those species the female of C. opulentus  spec. nov. is distinguished by a combination of atrium wide, occupying about 2/3 the width of the vulva and located on anterior third of the vulva ( Figs 30–31View FIGURES 30–32) (versus atrium much smaller, about 1/3 the width of the vulva, and usually located on the medial third of the vulva in C. penedentatus  [ Figs 46–47View FIGURES 46–51, 52–53View FIGURES 52–54] or atrium located on medial third of the vulva and wider, nearly as wide as the vulva, in C. ubicki  spec. nov. [ Figs 74–75, 77–78View FIGURES 74–79]); copulatory ducts and spermathecal stalks short and spermathecal stalks noncontiguous along midline of vulva ( Figs 31–32View FIGURES 30–32) (versus copulatory ducts and spermathecal stalks relatively long and spermathecal stalks contiguous along vulval midline in C. simplex  [ Figs 65–67View FIGURES 64–67]; copulatory ducts contiguous at the atrium as well as partway between the atrium and the spermathecal heads ( Figs 31–32View FIGURES 30–32) (versus contiguous only at the atrium in C. pan  spec. nov. [ Figs 38–39View FIGURES 37–41] and C. vulpinus  [ Figs 40–41View FIGURES 37–41]).

Description. Ventral tibia I macrosetae 2-1p-2-1p-1p(or 2 or 0).

Male: (n=23). Patellar apophysis ( Fig. 27View FIGURES 27–29) relatively short, length about 2/3 width of patella. Tip of proximal arm of tegular apophysis ( Figs 28View FIGURES 27–29, 55View FIGURES 55–58) elongate. Measurements (n=15). CL 2.43–3.3 (2.9±0.2), CW 1.80–2.35 (2.09±0.17), SL 1.25–1.58 (1.40±0.09), SW 1.17–1.45 (1.34±0.08). Holotype CL 2.8, CW 1.93, SL 1.33, SW 1.30.

Female: (n=20).Atrium ( Fig. 30View FIGURES 30–32) transverse, arched. Copulatory ducts ( Figs 31–32View FIGURES 30–32) broad at atrium, convergent anteriorly along midline of vulva then turning posteriorly. Measurements (n =18). CL 2.40–3.2 (2.8±0.2), CW 1.60–2.18 (1.94±0.17), SL 1.18–1.50 (1.36±0.09), SW 1.09–1.45 (1.30±0.09).

Distribution and natural history. South central Sierra Nevada region of eastern California south and west of Lake Tahoe in Alpine, El Dorado, and Placer Counties ( Fig. 86View FIGURE 86). Mature males have been collected from August through September and in December.


Royal British Columbia Museum - Herbarium


American Museum of Natural History


California Academy of Sciences


Tavera, Department of Geology and Geophysics


Departamento de Geologia, Universidad de Chile