Cybaeus consocius Chamberlin and Ivie,

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade, Zootaxa 4965 (3), pp. 401-436: 404-406

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Cybaeus consocius Chamberlin and Ivie


Cybaeus consocius Chamberlin and Ivie 

Figs 2–9View FIGURES 1–4View FIGURES 5–9, 16–17View FIGURES 13–17, 20View FIGURES 20–22, 84View FIGURE 84

Cybaeus consocius Chamberlin and Ivie 1932: 25  , fig. 61. Roewer 1954: 90. Bonnet 1956: 1301. Roth and Brown 1986: 3. Bennett 2006: 481, figs 23–25. Copley et al. 2009: 372, fig. 7. World Spider Catalog 2021.

Cybaeus marinensis Chamberlin and Ivie 1932: 25  , fig. 60. Roewer 1954: 91. Bonnet 1956: 1302.

Cybaeus angelus Chamberlin and Ivie 1942: 16  , fig. 50. Roewer 1954: 9. Roth and Brown 1986: 3.

Cybaeus mariensis: Roth and Brown 1986: 4  (incorrect subsequent spelling of C. marinensis  ).

Type material examined. U.S.A.: California: Holotype female of C. consocius, Marin County  , no date, R  . V  . Chamberlin ( AMNH), subsequently lost  . Neotype male of C. consocius  , designated by Bennett (2006), Marin County, south end of S.P. Taylor State Park, 1.xi.1953, V  . D. Roth & G. Marsh ( AMNH)  . Holotype female of C. angelus  . Los Angeles County, Los Angeles , no date, no collector ( AMNH)  . Holotype female of C. marinensis  . Marin County, no date, R  . V  . Chamberlin ( AMNH)  .

See discussion of type material and synonymy in Bennett (2006).

Other material examined. U.S.A.: California: Marin. 1♀, E slope Mt Tamalpais, 14.v.1952, H.S. Dybas ( FMNH);  2♂ 3♀, Muir Woods, 5.ix.1927 ( AMNH);  1♀, Muir Woods , 24.x.1953, V.D. Roth ( CAS);  1♀, Ring Mountain Preserve , S of Corte Madera, 18.xii.1982, V.F. Lee ( CAS);  3♀, Ross Bald Hill , 1.iii.1991, D. Ubick & T. Briggs ( CAS);  1♀ San Anselmo, San Rafael Ridge, 19.ix.1995, L.G. Freihofer ( CAS);  Mendocino. 4♀, near Lake Mendocino , P. R. Craig & C. Kirsch ( CAS);  Napa. 3♀, 2mi. W Oakville, 31.xii.1953 ( AMNH);  1♀, 3 mi W Oakville , 15.ii.1954, V.D. Roth & R. O. Schuster, ( AMNH);  1♀, 7 mi. W Oakville, 15.ii.1954 ( AMNH);  2♀, 7 mi. W Oakville , 13.xii.1957, L.M. Smith & R.O. Schuster ( CAS);  3♀, Skyline Park, Camp Coombs , 22.i.1999, T. Briggs, W. Rauscher & D. Ubick ( CAS);  San Francisco. 1♀, San Francisco, Marx ( MCZ);  Santa Clara. 2♀, Alum Rock Park , 22.ii.2013, I.M. Sokolov ( CAS);  Solano. 1♀, Cordelia , 14.9 mi N of Benicia, 9.ii.1964, P. R. Craig & D.L. Craig ( CAS)  .

Diagnosis. The male of C. consocius  is unlikely to be confused with the males of the other species in the consocius  group and is diagnosed by the unique forked tip of the proximal arm of the tegular apophysis ( Figs 3–4View FIGURES 1–4, 20View FIGURES 20–22). In ventral view the two forks are at right angles to each other with the larger, ventral-most fork directed prolaterally and featuring a sharp-edged longitudinal ridge medially along the ventral surface while the smaller fork is directed proximally and resembles the keel of a sailboat. The female of C. consocius  is only likely to be confused with the females of C. hesper  or C. hummeli  spec. nov.; those three species have similar vulval morphology featuring, in dorsal view, a distinctive U-shaped proximal loop of each copulatory duct ( Figs 7View FIGURES 5–9, 15View FIGURES 13–17, 25View FIGURES 23–26). In C. consocius  and C. hesper  the atrium is undivided and of an inverted U-shaped form ( Figs 5–14, 16–17View FIGURES 5–9View FIGURES 10–12View FIGURES 13–17); in C. hummeli  spec. nov. the atrium is divided into a pair of longitudinal slits or depressions separated by a medial convex septum ( Figs 23–24, 26View FIGURES 23–26). The females of C. consocius  and C. hesper  can be separated by differences in atrial and copulatory duct morphology. The medial area of the atrium between the lateral arms of the inverted “U” in C. consocius  is smoothly concave ( Figs 5, 9View FIGURES 5–9, 16View FIGURES 13–17) versus laterally concave and medially convex ( Fig. 13View FIGURES 13–17) in C. hesper  . The copulatory ducts in C. consocius  are attached to the lateral margins of the atrium and extend at best only slightly anterior of the anterior margin of the atrium ( Figs 6–8View FIGURES 5–9, 17View FIGURES 13–17); in C. hesper  the copulatory ducts are attached to the anterolateral margins of the atrium and extend well anterior of the anterior margin of the atrium ( Figs 14–15View FIGURES 13–17).

Description. Ventral tibia I macrosetae: 2-1p-2-1p-0.

Male: (n=3). Length of patellar apophysis about 3/4 width of patella, six small peg setae dorsally and around tip ( Fig. 2View FIGURES 1–4). Measurements (n=3). CL 2.20, 2.25, 2.6; CW 1.58, 1.68, 1.85; SL 1.11, 1.13, 1.27; SW 1.05, 1.07, 1.21. Neotype is largest specimen.

Female: (n=31). Copulatory ducts ( Figs 6–8View FIGURES 5–9, 17View FIGURES 13–17) somewhat membranous, contiguous or separate at atrium. Spermathecal stalks well separated ( Fig. 7View FIGURES 5–9). Measurements (n=14). CL 1.60–2.43 (2.11±0.23), CW 1.09–1.57 (1.44±0.15), SL 0.88–1.17 (1.08±0.08), SW 0.81–1.10 (1.01±0.08). Holotypes: C. consocius CL  2.20, CW 1.53, SL 1.12, SW 1.04; C. angelus CL  2.30, CW 1.57, SL 1.13, SW 1.04; C. marinensis CL  2.43, CW 1.70, SL 1.17, SW 1.10.

Distribution and natural history. Coastal California from southern Mendocino County to northern Santa Clara County ( Fig. 84View FIGURE 84). Most records are from the San Francisco Bay area. Following Bennett (2006), we believe the record from the Los Angeles area (a single specimen, the holotype of C. angelus  ) to be erroneous: no other species of the consocius  group is known to range that far south and no further specimens of C. consocius  have been recorded from the Los Angeles area subsequent to the original description of C. angelus  . This record is not included on the distribution map. Also see distribution discussion under C. hesper  . Mature males have been collected in September and November.


Departamento de Geologia, Universidad de Chile


Royal British Columbia Museum - Herbarium


American Museum of Natural History


Field Museum of Natural History


California Academy of Sciences


Tavera, Department of Geology and Geophysics


Museum of Comparative Zoology














Cybaeus consocius Chamberlin and Ivie

Bennett, Robb, Copley, Claudia & Copley, Darren 2021

Cybaeus mariensis:

Roth, V. D. & Brown, W. L. 1986: 4

Cybaeus angelus

Roth, V. D. & Brown, W. L. 1986: 3
Roewer, C. F. 1954: 9
Chamberlin, R. V. & Ivie, W. 1942: 16

Cybaeus consocius

Copley, C. R. & Bennett, R. & Perlman, S. J. 2009: 372
Bennett, R. G. 2006: 481
Roth, V. D. & Brown, W. L. 1986: 3
Bonnet, P. 1956: 1301
Roewer, C. F. 1954: 90
Chamberlin, R. V. & Ivie, W. 1932: 25

Cybaeus marinensis

Bonnet, P. 1956: 1302
Roewer, C. F. 1954: 91
Chamberlin, R. V. & Ivie, W. 1932: 25