Cochlostoma (Clessiniella) waldemari ( Wagner, 1897 )

Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán & Gittenberger, Edmund, 2021, Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 762 (1), pp. 49-95: 87-89

publication ID

publication LSID


persistent identifier

treatment provided by


scientific name

Cochlostoma (Clessiniella) waldemari ( Wagner, 1897 )


Cochlostoma (Clessiniella) waldemari ( Wagner, 1897)  

Figs 1 View Fig , 3D View Fig , 14E View Fig , 15E View Fig , 16E View Fig , 31–33 View Fig View Fig View Fig , 34 View Fig (top), 35 (top), 36 (bottom)

Pomatias (Auritus) waldemari A.J. Wagner, 1897: 616   , pl. 9, 90 (“Ogulin”).

Cochlostoma waldemari   – Welter-Schultes 2012: 103.


Spotless shell. Moderately strong, more or less widely spaced ribs. Columellar lobe gently curved backwards. External lobe of type “II” in almost all the populations. The body spermiduct is huge and twisted instead of straight as in all the other species of Clessiniella   .

Type material

Syntypes CROATIA • ♀; Ogulin; A.J. Wagner leg.; MIZPAS-W-6979a ( Fig. 33 View Fig )   2 ♀♀, 3 ♂♂; same collection data as for preceding; MIZPAS-W-6979/5   .

Other material

CROATIA • Kapela pass , Ogulin; 45.07534° N, 15.20837° E; 2009; De Mattia leg.; EZ0909 GoogleMaps   Klek Mt , close to the top; 45.25904° N, 15.14044° E; 2009; Zallot leg.; EZ0904 GoogleMaps   Klek Mt , cliffs halfway ; 45.25438° N, 15.14561° E; 1996; Zallot leg.; EZ0905 GoogleMaps   Mt Risnjak ; 45.43095° N, 14.6188° E; 1400 m a.s.l.; 2009; De Mattia leg.; EZ0845 GoogleMaps   Mt Risnjak ; 45.39072° N, 14.61748° E; 1000 m a.s.l.; 2010; Zallot leg.; EZ0907 GoogleMaps   Ročko Polje ; 45.3607° N, 14.09534° E; 2009; De Mattia leg.; WdM6697 GoogleMaps   Rudnica ; 45.23402° N, 15.33827° E; 2009; De Mattia leg.; EZ0906 GoogleMaps   Spilja Vrelo ; 45.31777° N, 14.71065° E; 2009; Zallot leg.; EZ0899 GoogleMaps   Vitunj-Ogulin ; 45.29114° N, 15.14041° E; 2005; Zallot leg.; EZ0903 GoogleMaps   .

SLOVENIA • Benete , 45.76192°N, 14.56752°E;2009; De Mattia leg.; WdM6805 GoogleMaps   Bresovica ; 45.95607°N, 14.43533° E; 2006; De Mattia leg.; EZ0897 GoogleMaps   Bresovica pri Predgradu ; 45.55164° N, 15.0437° E; 2009; De Mattia leg.; WdM6612 GoogleMaps   Divje Jezero ; 45.9827° N, 14.0277° E; 2009; De Mattia leg.; WdM6840 GoogleMaps   Dol-E of Čepovan ; 46.06445° N, 13.81189° E; 2009; De Mattia leg.; WdM6837 GoogleMaps   Jelenja Vas ; 45.5115° N, 15.05346° E; 2009; De Mattia leg.; WdM6804 GoogleMaps   Krempa Kolpa ; 45.53417° N, 15.09528° E; 1996; Mildner leg.; EZ0902 GoogleMaps   Marija Ceplje Kolpa ; 45.53444° N, 15.09389° E; 2006; Mildner leg.; EZ0901 GoogleMaps   Mt Snežnik 1150; 45.56913° N, 14.321° E; Jun. 2010; De Mattia leg.; WdM7014 GoogleMaps   Mt Snežnik 1620; 45.58845° N, 14.44209° E; Jun. 2010; De Mattia leg.; EZ0963 GoogleMaps   Mt Snežnik car parking ; 45.58235° N, 14.43117° E; Jun. 2010; De Mattia leg.; WdM7006 GoogleMaps   Polog-E of Čepovan ; 46.06387° N, 13.81275° E; 2009; De Mattia leg.; WdM6842 GoogleMaps   SW Pescena Glava; 46.05156° N, 13.73844° E; 843 m a.s.l.; 2009; De Mattia leg.; WdM6829 GoogleMaps   Tisovec ; 45.81809° N, 14.76859° E; 2009; De Mattia leg.; WdM6749 GoogleMaps   same collection data as for preceding; WdM6750 GoogleMaps   Vrsic Pass ; 46.44278° N, 13.76417° E; 1985; Zallot leg.; EZ0896 GoogleMaps   Zagradec- Polyane ; 45.86028° N, 14.83417° E; Bodon leg.; EZ0900 GoogleMaps   .


SHELL. The shell is more or less light corneous brown and spotless. The size of the aperture varies considerably among the populations; shells with a large aperture and a correspondingly large lip occur next to shells with a poorly developed lip and a relatively small aperture. The lip is gently curved backwards on the columellar side.

FEMALE GENITALIA. The female genitalia are hardly distinguishable from those of Co. (Cl.) tergestinum   if not for the more relaxed loops.

MALE GENITALIA. There is a long, inflated penis. The penial spermiduct is slightly twisted and occupies only the central part of the penis. The body spermiduct is twisted and conspicuously protruding (see Figs 34–35 View Fig View Fig ).


The species inhabits the north-western and south-eastern part of Slovenia and an adjacent area in Croatia. It is reported from Austria for the Karawanken and the Gailtaler Alpen ( Edlinger & Mildner 1979). Its range largely surrounds that of Co. (Cl.) tergestinum   (see Fig. 1 View Fig ).

Habitat preference

The snails are found on rocky cliffs as well as on boulders in woody areas.


To assign a given population to Co. (Cl.) waldemari   rather than to Co. (Cl.) tergestinum   on the basis of the morphology of the shell and the female genitalia alone could be challenging if not impossible (see Figs 34–35 View Fig View Fig ). Only the structure of the male genitalia allows a convincing identification of the species. The H3 analysis ( Fig. 7 View Fig ) confirms that these sibling species belong to separate clades within Clessiniella   .

Boeckel (1939) considered Co. (Cl.) waldemari   a subspecies of Co. (Cl.) tergestinum   . This view was contradicted by Bole (1994). In some areas, like on Mount Snežnik in southern Slovenia, the two species live not far from each other and here they can be distinguished also conchologically while differing in the presence or absence of the spots on the teleoconch (with Co. (Cl.) waldemari   having a spotless shell) and in the shape of the body whorl approaching the aperture, in lateral view ( Fig. 36 View Fig ).














Cochlostoma (Clessiniella) waldemari ( Wagner, 1897 )

Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán & Gittenberger, Edmund 2021

Cochlostoma waldemari

Welter-Schultes F. W. 2012: 103

Pomatias (Auritus) waldemari A.J. Wagner, 1897: 616

Wagner A. J. 1897: 616