Cochlostoma (Clessiniella) anomphale Boeckel, 1939

Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán & Gittenberger, Edmund, 2021, Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 762 (1), pp. 49-95 : 75-78

Cochlostoma (Clessiniella) anomphale Boeckel, 1939

Figs 1 , 3A, 3A 1 , 14A , 15A , 16A , 22–24

Cochlostoma septemspirale anomphale Boeckel, 1939: 41–50 (“Karawanken, Trögern. Weg nach Vellach”).

Cochlostoma septemspirale anomphale – Zilch 1958: 57.

Cochlostoma (Turritus) anomphale – Gittenberger 1971: 115–118.

Cochlostoma anomphale – Welter-Schultes 2012: 91.

Diagnosis

Shell quite densely ribbed, spots on the teleoconch generally clearly visible, body whorl inflated shortly before the lip. Visceral oviduct simplified with 1–2 superficial loops at most; seminal receptacle always club-shaped.

Type material

Paratypes AUSTRIA • 2 specs; Trögern , way to Vellach; SMF 42967/2 ( Fig. 24 ) .

Other material (we analyzed fresh material from the type locality)

AUSTRIA • EisenKappel ; 46.48583° N, 14.59111° E; 2010, Mildner leg.; EZ0034 • Vellach ; 46.42332° N, 14.53639° E; Aug. 2010; De Mattia leg.; EZ0991 .

SLOVENIA • 210 road, N of Kokra; 46.34318° N, 14.48336° E; Aug. 2010; De Mattia leg.; EZ0993 • Gornji Grad; 46.2934° N, 14.82796° E; Aug. 2010; De Mattia leg.; EZ0996 • Kamniska Bistrika ; 46.32722° N, 14.59° E; 2010; Zallot leg.; EZ0033 • Kokra ; 46.30356° N, 14.4771° E; Aug. 2010; De Mattia leg.; EZ0995 • Logarska Dolina ; 46.41959° N, 14.65074° E; Aug. 2010; De Mattia leg.; EZ0994 • Logarska Dolina ; 46.42334° N, 14.66301° E; Aug. 2010; De Mattia leg.; EZ0992 • Podvolovliek- Stahovica ; 46.28787° N, 14.69765° E; Aug. 2005; Zallot leg.; EZ0036 • Savinjska Dolina ; 46.36596° N, 14.7318° E; 2010; De Mattia leg.; WdM1579 • Solčava; 46.42002° N, 14.19172° E; Jul. 2010; De Mattia leg.; WdM7010 • Stahovika ; 46.26528° N, 14.60361° E; 2005; Zallot leg.; EZ0035 • SV Anton ; 46.28787° N, 14.69765° E; 2010; Zallot leg.; EZ0032 .

Remarks

The holotype and 23 paratypes should be in the Boeckel collection at the Naturkundemuseum Erfurt ( Hartmann et al. 2018: 26). However, the shells could not be traced when this article was prepared (C. Unger in mail October 14, 2020).Type locality - Austria, Trögern, way to Vellach.

Description

SHELL. The outside of the body whorl is inflated shortly before the apertural lip ( Fig. 4A : OB). The lip is poorly developed; it is almost vertical at the palatal side, and curved backwards and covering the umbilicus at the columellar side. The shells are more densely sculptured and the ribs are weaker than in the other species of Clessiniella . Most of the shells have spots.

FEMALE GENITALIA. There is a spindle-like seminal receptacle. The visceral oviduct has only 1–2, superficial, loops.

MALE GENITALIA. The swollen penis is longer than the body and has a central spermiduct. There is a protruding, straight, body spermiduct.

Distribution

Cochlostoma (Clessiniella) anomphale inhabits the Karawanken in south-eastern Austria and northern Slovenia.

Habitat preference

The animals can be found under and on stones and boulders in shady habitats in forests.

Remarks

This species was initially described as a subspecies of Cochlostoma (Cochlostoma) septemspirale . Gittenberger (1971) considered it a separate species because of the sympatry with Co. (Co.) semptemspirale heydenianum and the lack of conchologically intermediate specimens. This is an epitome of quite common Cochlostoma taxonomical issues: based on shell morphology it was initially described as subspecies of a taxon which is only distantly related.

Figures

Fig. 1. Map of the sampling localities. Blue dots = Cochlostoma (Clessiniella) villae (Strobel, 1851); red dots = Co. (Cl.) stelucarum sp. nov.; green dots = Co. (Cl.) tergestinum (Westerlund, 1878); yellow dots = Co. (Cl.) waldemari (A.J. Wagner, 1897); orange dots = Co. (Cl.) anomphale Boeckel, 1939.

Fig. 3. Examples of protoconchs: the red dots mark the end of the protoconchs. A. Cochlostoma (Clessiniella) anomphale Boeckel, 1939 (EZ0034). A1. Co. (Cl.) anomphale (EZ0034): 1 = end of the first smooth part of the protoconch; 2 = end of the protoconch. B. Co. (Cl.) tergestinum (Westerlund, 1878) (EZ0799). C. Co. (Cl.) villae (Strobel, 1851) (EZ0885). D. Co. (Cl.) waldemari (A.J. Wagner, 1897) (WdM6697). E. Co. (Cl.) stelucarum sp. nov. (EZ0104).

Fig. 14. The five species of Clessiniella Zallot et al., 2015. A–D. Picture of a female (top) and male (bottom) specimen from the type localities. A. Cochlostoma (Clessiniella) anomphale Boeckel, 1939. Vellach, Austria (EZ0991). B. Co. (Cl.) stelucarum sp. nov., Cornino, Italy (EZ0104). C. Co. (Cl.) tergestinum (Westerlund, 1878), Val Rosandra, Italy (EZ0840). D. Co. (Cl.) villae (Strobel, 1851), Mizzole, Italy (EZ0878). E. Specimens from a locality close to the type one: Co. (Cl.) waldemari (A.J.Wagner, 1897), Mt Klek, Croatia (EZ0905).

Fig. 15.Comparison of femalegenitalia.A. Cochlostoma (Clessiniella) anomphaleBoeckel, 1939.Vellach, Austria (EZ0991). B. Co. (Cl.) stelucarum sp. nov., Cornino, Italy (EZ0104). C. Co. (Cl.) tergestinum (Westerlund, 1878), Val Rosandra, Italy (EZ0840). D. Co. (Cl.) villae (Strobel, 1851), Mizzole, Italy (EZ0878). E. Co. (Cl.) waldemari (A.J.Wagner, 1897), Mt Klek, Croatia (EZ0905).

Fig. 16.Comparisonof male genitalia.A. Cochlostoma (Clessiniella)anomphaleBoeckel, 1939.Vellach, Austria (EZ0991). B. Co. (Cl.) stelucarum sp. nov., Cornino, Italy (EZ0104). C. Co. (Cl.) tergestinum (Westerlund, 1878), Val Rosandra, Italy (EZ0840). D. Co. (Cl.) villae (Strobel, 1851), Mizzole, Italy (EZ0878). E. Co. (Cl.) waldemari (A.J.Wagner, 1897), Tisovec, Croatia (WdM6750).

Fig. 22.Shell variability in Cochlostoma (Clessiniella) anomphale Boeckel, 1939.A. Savinjska Dolina, Slovenia (WdM1579). B. Eisenkappel, Austria (EZ0034).C. Stahovika, Slovenia (EZ0035).D. Vellach, Austria (EZ0991). E. Solčava, Slovenia (WdM7010). F. Logarska Dolina, Slovenia (EZ0994).

Fig. 23. Variation of the female genitalia in Cochlostoma (Clesssiniella) anomphale Boeckel, 1939. A. Eisenkappel, Austria (EZ0034). B. Savinjska Dolina, Slovenia (WdM1579). C. Vellach, Austria (EZ0991). D. Logarska Dolina, Slovenia (EZ0994). E1–E2. Solčava, Slovenia (WdM7010).

Fig. 24. Cochlostoma (Clessiniella) anomphale Boeckel, 1939. Paratypes at the Senckenberg Museum Frankfurt (SMF42967/2).

Fig. 4. Features of the shell of Clessiniella Zallot et al., 2015. A. Shape of the body whorl approaching the aperture. B. Strength of the spots on the shell (0 = spotless shell to 2 = really evident spots). C. Strength of the ribs (1 = weak ribs to 3 = really prominent ribs). D. Strength of the ribs on the external side of the body whorl approaching the aperture (W= smooth shell; I = intermediate; S = prominent ribs).