Orthochirus scrobiculosus ( Grube, 1873 ), 1993

Kovařík, Baloorthochirus becvari & 1996, 2004, Revision and taxonomic position of genera Afghanorthochirus Lourenço & Vachon, Baloorthochirus Kova ík, Butheolus Simon, Nanobuthus Pocock, Orthochiroides Kova ík, Pakistanorthochirus Lourenço, and Asian Orthochirus Karsch, with descriptions of twelve new species (Scorpiones, Buthidae), Euscorpius 16, pp. 1-33 : 19-20

publication ID

1536-9307

publication LSID

lsid:zoobank.org:pub:4D38E32B-A445-4707-ABEE-3C6D3BFA4A2E

persistent identifier

https://treatment.plazi.org/id/038B2040-3C24-5B3F-FF77-FDB8FA97FC61

treatment provided by

Carolina

scientific name

Orthochirus scrobiculosus ( Grube, 1873 )
status

 

Orthochirus scrobiculosus ( Grube, 1873)

Androctonus scrobiculosus Grube, 1873: 56 .

Butheolus melanurus: Kraepelin, 1899: 35 ; Pocock, 1900: 28; Birula, 1900: 13; Birula, 1900: 373; Kraepelin, 1901: 267; Kraepelin, 1903: 566; Birula, 1903: 74; Birula, 1904: 32; Birula, 1905: 125; Schenkel, 1932: 381; Caius, 1938: 579;? Moriggi, 1941: 90; Hadži, 1943: 124;? Bücherl, 1959: 257;? Pérez Minnocci, 1974: 20.

Orthochirus melanurus: Kraepelin, 1895: 84 ; Birula, 1898: 281; Roewer, 1943: 208;? Tikader & Bastawade, 1983: 134.

Orthochirus melanurus forma typica: Birula, 1900: 13.

Butheolus scrobiculosus: Birula, 1909: 356 ; Birula, 1911: 174; Lampe, 1918: 194.

Butheolus scrobiculatus: Roewer, 1943: 208 .

Orthochirus scrobiculosus: Birula, 1917: 215 ; Vachon, 1940: 181; Whittick, 1947: 124 (in part); Vachon, 1949: 136 (1952: 222); Vachon, 1959: 166;? Khalaf, 1962: 2;? Khalaf, 1963: 64; Habibi, 1971: 44; Farzanpay & Pretzmann, 1974: 216; Pérez Minnocci, 1974: 28; Fet, 1980: 225; Fet, 1981: 168; Levy & Amitai, 1980: 94; Farzanpay, 1988: 40; Fet, 1989: 113; Polis & Sissom, 1990: 185; Simard & Watt, 1990: 421; Sissom, 1990: 92; El-Hennawy, 1992: 129; Kovařík, 1993: 203; Amr & El-Oran, 1994: 187; Fet, 1994: 529; Lourenço, 1997: 153; Kovařík, 1998: 117; Kovařík, 1998: 116; Fet & Lowe, 2000: 196; Soleglad & Fet, 2003: 5.

Orthochirus scrobiculosa :? Pringle, 1960: 78.

Butheolus scrobiculosus scrobiculosus: Birula, 1909: 359 .

Orthochirus scrobiculosus scrobiculosus: Birula, 1917: 241 ; Birula, 1928: 83; Pérez Minnocci, 1974: 28; Levy & Amitai, 1980: 94; Fet, 1989: 116; Fet, 1994: 529; Kovařík, 1997: 49; Fet & Lowe, 2000: 197.

= Androctonus melanurus Kessler, 1874: 16 (TL: Novoaleksandrovskoye, Mangyshlak Region, now Kazakstan; ZISP) (syn. by Birula, 1909: 358).

Butheolus scrobiculosus melanurus: Birula, 1909: 359 .

Orthochirus scrobiculosus melanurus: Birula, 1917: 241 ; Birula, 1918: 40; Birula, 1928: 83; Vachon, 1959: 169; Vachon, 1966: 214; Levy & Amitai, 1980: 94; Fet, 1989: 117; Fet, 1994: 529; Kovařík, 1997: 49; Kovařík, 1998: 116; Fet & Lowe, 2000: 198.

Orthochirus melanurus forma beta (intermedia): Birula, 1898: 282.

Butheolus melanurus typicus Birula, 1900: 373 ;? Pocock, 1900: 29.

= Buthus schneideri L. Koch, 1878: 61 (TL: Krasnowodsk; MNHN) (syn. by Pocock, 1890: 121).

Butheolus schneideri: Simon, 1889b: 386 .

Orthodactylus schneideri: Karsch, 1886: 76 (in part); Pocock, 1889: 117; Kraepelin, 1891: 215 (in part).

= Orthodactylus olivaceus Karsch, 1881: 91 (TL: Sicily (incorrect type locality); ZMHB); Moritz & Fischer, 1980: 321 (syn. by Karsch, 1886: 76)

Orthochirus olivaceus: Birula, 1928: 83 ; Vachon, 1959: 166.

= Butheolus conchini Simon, 1889b: 386 (TL: Bely- Bugor [=Aktepe], Turkmenistan; MNHN); Kraepelin, 1895: 84; Kraepelin, 1899: 36; Birula, 1905: 126; Roewer, 1943: 208; Pérez Minnocci, 1974: 20; (syn. by Birula, 1898: 281).

Butheolus melanurus conchini: Birula, 1900: 374 ; Birula, 1904: 32; Birula, 1905: 128.

Orthochirus conchini: Levy & Amitai, 1980: 94 .

=? Orthochirus melanurus forma gama (concolor) Birula, 1898: 282 (TL: Karshi, Uzbekistan; ZISP).

Butheolus melanurus concolor: Birula, 1900: 374 ; Birula, 1904: 32.

Butheolus scrobiculosus concolor: Birula, 1909: 359 ; Werner, 1916: 81; Birula, 1918: 40; Lampe, 1918: 194.

Orthochirus scrobiculosus concolor: Birula, 1917: 241 ; Birula, 1918: 40; Vachon, 1959: 169; Pérez Minnocci, 1974: 29; Levy & Amitai, 1980: 94; Fet, 1989: 118; Fet, 1994: 530; Kovařík, 1998: 116; Fet & Lowe, 2000: 198.

=? Butheolus melanurus dentatus Birula, 1900: 375 ; Pérez Minnocci, 1974: 20 (Husseinabad, Seistan, Persia; ZISP).

Orthochirus scrobiculosus dentatus: Fet, 1989: 116 ; Fet, 1994: 530; Kovařík, 1997: 49; Kovařík, 1998: 116; Fet & Lowe, 2000: 198.

=? Orthochirus scrobiculosus mesopotamicus Birula, 1918: 35 (TL: Mesopotamia inferior, Iraq; ZISP); Birula, 1928: 83; Vachon, 1966: 214; Pérez Minnocci, 1974: 28; Levy & Amitai, 1980: 94; Fet, 1989: 116; El-Hennawy, 1992: 130; Fet, 1994: 530; Kovařík, 1998: 116; Fet & Lowe, 2000: 199.

=? Butheolus melanurus persa Birula, 1900: 374 (TL: Seistan, Kirman, Birdschan, Iran; ZISP); Birula, 1903: 75; Birula, 1905: 145; Weidner, 1959: 99; Pérez Minnocci, 1974: 20.

Butheolus scrobiculosus persa: Birula, 1909: 359 ; Penther, 1912: 114; Birula, 1918: 35.

Orthochirus scrobiculosus persa: Birula, 1917: 241 ; Birula, 1918: 40; Birula, 1928: 83; Levy & Amitai, 1980: 94; Fet, 1989: 116; Fet, 1994: 530 (statut?); Kovařík, 1997: 49; Kovařík, 1998: 116; Fet & Lowe, 2000: 199.

Orthochirus persa: Vachon, 1966: 213 ; Habibi, 1971: 44; El-Hennawy, 1992: 129.

TYPE LOCALITY AND TYPE REPOSITORY. Lenkoran (incorrect type locality; see Fet & Lowe, 2000: 196); UWCP.

TYPE MATERIAL EXAMINED. Italy, Sicily (incorrect type locality), 1♂ A (holotype of Orthodactylus olivaceus Karsch, 1881 ), leg. Schneider, ZMHB No. 3629 .

OTHER MATERIAL EXAMINED. Afghanistan, Prov. Mazar-i-Sharif , 9.XII–17.I. 1964, 365 m (loc. No. J. 2), 1♂ E, leg. Jakeš, MMBC , for locality details see Jakeš & Povolný (1967); Old Balkh, 6.V.1970, 1♂ 2♀ A, leg. E. S. Ross, CASC ; 20 km SW Dosi , 7.V.1970, 1♂ (im.)A, leg. E. S. Ross, CASC . Iran, 10 km W of Baqbaqú (Khorásán), 36°05’N 60°25’E, 10.V.1997, 680 (–) m, 2♀ A, leg. M. Kaftan, FKCP GoogleMaps . Turkmenistan, Ashkhabad , 27.IV.1987, 1♂ E, leg. P. Slabý, FKCP ; Ashkhabad , 1988, 1♀ E, leg. Šeda, FKCP ; V.1988, 1♀ E, leg. M. Král, FKCP ; South Ashkhabad , V.1990, 2♀ E, leg. Podhajský, FKCP ; Ashgabat, Bagir , 800 m, 16.V.1996, 1♂ E, leg. Liehtfang, FKCP ; Badkhyz Nature Reserve, ca. 14–16 km SSW of Kyzyldzhar cordon, near Eroilanduz Salt Lake , 340–350 m, (35º40'33"N 61º49'18"E - 35º42'04"N 61º48'53"E), 1♂ 2♀ A, leg. A. V. Gromov, FKCP GoogleMaps . Uzbekistan, Derbent , 2.V.1981, 1juv. E, leg. S. Bečvář, FKCP ; Muratan Range , V.1989, 1♂ 1♀ 1juv. E, leg. D. Král, FKCP ; Bukhara, Djeiran Pitomnik ( Gazelle Nursery ), VI.1989, 1♂ 1♀ E, leg. J. Růžička, FKCP ; Nuratinski Reserve, Chajatsaj , 1400 m, 15.V.1996, 1♀ A, FKCP .

DIAGNOSIS: Fourth and fifth metasomal segments ventrally punctate and without carinae. Spaces among punctae smooth, without granules. Dorsal surface of all metasomal segments mesially smooth, without granules. Entire metasoma glabrous (very short, thin setae may issue from punctae). Mesosoma and metasoma black, telson reddish brown, legs yellow, pedipalps yellow to greenish yellow. Movable fingers of pedipalps bear 8–10 rows of granules with internal granules and without external granules. Movable fingers with 2–5 distal granules. Tarsus of first to third legs with bristlecombs, fourth without. Pectinal teeth number 15–18 in females and 19–20 in males.

DISTRIBUTION. Afghanistan (Pocock, 1889: 118), Iran (Vachon, 1966: 214; Habibi, 1971: 44), Kazakhstan ( Fet, 1989: 117), Tajikistan ( Fet, 1989: 117), Turkmenistan (Simon, 1889b: 386), and Uzbekistan ( Fet, 1989: 118).

Published records for Djibouti (Kraepelin, 1901: 267), Egypt (Kraepelin, 1899: 36), India (Pérez Minnocci, 1974: 28; Tikader & Bastawade, 1983: 140), Iraq ( Khalaf, 1962: 2), Israel ( Bücherl, 1959: 257), Pakistan (Kraepelin, 1899: 36; Pérez Minnocci, 1974: 28), and Somalia (Moriggi, 1941: 90) must be regarded as erroneous.

COMMENTS: Despite several attempts, it has unfortunately not been possible to examine the types of taxa described by Birula, which have hitherto been regarded as subspecies of O. scrobiculosus . It concerns O. s. concolor Birula, 1898, O. s. dentatus (Birula, 1900), O. s. mesopotamicus Birula, 1918, and O. s. persa (Birula, 1900) . In order to complete the revision, I thus have no choice but to leave the taxonomic positions of those taxa unresolved, and hope that the situation can be corrected in a near future. Birula (1918: 39–40) clearly characterized O. s. concolor Birula, 1898 as lacking external granules on movable fingers, and O. s. mesopotamicus ( Iraq) and O. s. persa ( Iran, Pakistan) as possessing these granules. It is therefore very likely that these two taxa in reality are separate species of which O. s. mesopotamicus is close to O. iraqus sp. n., which may even prove to be its junior synonym. Similarly, one of the four new species described here from Iran could prove to be a junior synonym of O. s. persa .

UWCP

University of Wroclaw

MMBC

Moravske Muzeum [Moravian Museum]

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Scorpiones

Family

Buthidae

Genus

Orthochirus

Loc

Orthochirus scrobiculosus ( Grube, 1873 )

Kovařík, Baloorthochirus becvari & 1996 2004
2004
Loc

Orthochirus scrobiculosus dentatus:

KOVARIK 1997: 49
FET 1994: 530
FET 1989: 116
1989
Loc

Orthochirus persa:

HABIBI 1971: 44
1971
Loc

Orthochirus olivaceus:

BIRULA 1928: 83
1928
Loc

Orthochirus scrobiculosus: Birula, 1917: 215

FET 1994: 529
KOVARIK 1993: 203
FET 1989: 113
FARZANPAY 1988: 40
FET 1981: 168
FET 1980: 225
HABIBI 1971: 44
KHALAF 1963: 64
KHALAF 1962: 2
BIRULA 1917: 215
1917
Loc

Orthochirus scrobiculosus scrobiculosus:

KOVARIK 1997: 49
FET 1994: 529
FET 1989: 116
BIRULA 1928: 83
BIRULA 1917: 241
1917
Loc

Orthochirus scrobiculosus melanurus:

KOVARIK 1997: 49
FET 1994: 529
FET 1989: 117
BIRULA 1928: 83
BIRULA 1918: 40
BIRULA 1917: 241
1917
Loc

Orthochirus scrobiculosus concolor:

FET 1994: 530
FET 1989: 118
BIRULA 1918: 40
BIRULA 1917: 241
1917
Loc

Orthochirus scrobiculosus persa:

KOVARIK 1997: 49
FET 1994: 530
FET 1989: 116
BIRULA 1928: 83
BIRULA 1918: 40
BIRULA 1917: 241
1917
Loc

Butheolus scrobiculosus:

BIRULA 1911: 174
BIRULA 1909: 356
1909
Loc

Butheolus scrobiculosus scrobiculosus:

BIRULA 1909: 359
1909
Loc

Butheolus scrobiculosus melanurus:

BIRULA 1909: 359
1909
Loc

Butheolus scrobiculosus concolor:

BIRULA 1918: 40
BIRULA 1909: 359
1909
Loc

Butheolus scrobiculosus persa:

BIRULA 1918: 35
BIRULA 1909: 359
1909
Loc

Butheolus conchini

BIRULA 1905: 126
BIRULA 1898: 281
1905
Loc

Butheolus melanurus conchini:

BIRULA 1905: 128
BIRULA 1904: 32
1904
Loc

Butheolus melanurus concolor:

BIRULA 1904: 32
1904
Loc

Butheolus melanurus:

BUCHERL 1959: 257
HADZI 1943: 124
CAIUS 1938: 579
BIRULA 1905: 125
BIRULA 1904: 32
BIRULA 1903: 74
1903
Loc

Orthochirus melanurus

BIRULA 1898: 281
1898
Loc

Orthochirus melanurus

BIRULA 1898: 282
1898
Loc

Orthodactylus schneideri:

KARSCH 1886: 76
1886
Loc

Orthodactylus olivaceus

KARSCH 1886: 76
KARSCH 1881: 91
1881
Loc

Buthus schneideri L. Koch, 1878: 61

KOCH 1878: 61
1878
Loc

Androctonus melanurus

BIRULA 1909: 358
KESSLER 1874: 16
1874
Loc

Androctonus scrobiculosus

GRUBE 1873: 56
1873