Ulmus woodii, WHEELER AND, 2007

Wheeler, Elisabeth A., Manchester, Steven R. & Baas, Pieter, 2023, A late Eocene wood assemblage from the Crooked River Basin, Oregon, USA, PaleoBios 40 (14), pp. 1-55 : 22-23

CF. ULMUS WOODII WHEELER AND MANCHESTER, 2007

FIG. 11A–F

Description— Growth ring boundaries distinct (1p). Wood semi-ring-porous to ring-porous (3p, 4p). Latewood vessels in radial multiples and clusters (11p) arranged in wavy tangential bands (6p) ( Fig.11A–C ). Mean tangential diameters of earlywood vessels 70 ( SD =13) µm, range 48–87 µm. Perforation plates exclusively simple (13p, 14a). Intervessel pits crowded alternate (22p), polygonal in outline (23p), not vestured (29a), 7–11 µm in horizontal diameter (26p 27p); vessel-ray parenchyma pits similar in size to intervessel pits with borders somewhat reduced (31p) ( Fig. 11D ). Vessel element lengths 139–214 µm (n=7). Widely spaced thin-walled tyloses present.

Fibers non-septate (66p), pits not observed (62a).

Axial parenchyma paratracheal, non-crystalliferous strands usually 4 cells.

Rays mostly 3–5-seriate (98p), uniseriate rays not common ( Fig. 11E,F ); homocellular composed of procumbent cells (104p) ( Fig. 11D ); multiseriate ray heights average 267 µm ( SD =95), range 146–444 µm.

Solitary prismatic crystals in chambered axial parenchyma strands, not in ray parenchyma (136p 138a 142p) ( Fig. 11D,F ).

Pith composed of thin-walled isodiametric parenchyma.

Specimens— UF 278-62702, 84866, 84880, 84883, estimated maximum diameters of 4 cm, 2 cm with pith, 1.1 cm with pith.

Comments— Because of the tyloses, we could only

cells, Ab =absent, Occ =occasional; Ax Par =axial parenchyma distribution, Sc =scanty paratracheal, V =vasicentric, Al =aliform, Cf =confluent, Bnd =banded, M =marginal; Cry =crystals, AB =absent, RP =present in ray parenchyma, A =present in axial parenchyma;? =no information; * =small axis.

UF.278 WOODS POR V Grp V Arr IVP VRP RW Rcp ShC Ax Par Cry Sc, V, Urticaleoxylon stevensii / S-Rm Sim/ Ho, He SRP Ran 8–11 1–6 Ab Al, Cf, RP UF 278-84893 (2–3) Red (1) M S-Rms Sim/ 1–3 Ho, He Moroxylon ?/ UF 278-84899 RP Diag 8–11 Ab Cf, M Ab (Cl) Red (–4) (1–2)

Cannabaceae / Moraceae S-Rm

DP Ran Wood Type 1 UF 278-84894 (2–3)

Cannabaceae / Moraceae S-Rm

? * Ran Wood Type 2 UF 278-84906 (2–3)

Figures

Figure 11. Ulmaceae. Ulmus woodii. A. Pith composed of isodiameteric parenchymatous cells, first ring of wood with vessels mostly solitary and in radial alignment, TS, UF 278-84883. B. Rings 2 and 3 semi-ring-porous, TS, UF 278-84883. C. Outermost rings of the sample, latewood vessels in radial multiples and clusters in a ulmiform pattern, UF 278-84866. D‒F. Rays composed of procumbent cells, crowded alternate intervessel pits (IVP), vessel-ray parenchyma pits (VRP) of similar size to intervessel pits, crystalliferous axial parenchyma strands (C), RLS, UF 278-62702. E. Ray mostly 3‒5-seriate, uniseriate rays rare, TLS. F. Multiseriate rays, and crystalliferous axial parenchyma strands (C), TLS. Scale bars=200 µm in A–C; 100 µm in E; 50 µm in D, F.