Laccophilus schillhammeri, Hájek & Brancucci, 2015

Hájek, Jiří & Brancucci, Michel, 2015, A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae), Raffles Bulletin of Zoology 63, pp. 309-326 : 323-324

Laccophilus schillhammeri , new species

( Figs. 13 , 31 )

Type locality. Myanmar, Sagaing Region, Chatthin Wildlife Sanctuary, above San Myaung camp, ca. 23°36′N, 95°31′E.

Material examined. Holotype male, labelled: Myanmar (90A) Sagaing Division / Chatthin Wildlife Sanctuary / above San Myaung Camp, spring / 18.6.2002, leg. Schillhammer et al. ( NHMW) . Paratypes: 8 males, 7 females: same label data as holotype ( NHMB, NHMW, NMPC); 1 male: MYANMAR: Sagaing Division / Chatthin Wildlife Sanctuary / ca. 200m, 5.- 17.10.1998 / leg. Schillhammer (4) ( NHMW) ; 1 female: MYANMAR: Sagaing Division / Chatthin Wildlife Sanctuary / 23°31.481′N 95°38.804′E / ca. 260m, 9.10.1998 / leg. Schillhammer (9) ( NHMW) ; 2 females: MYANMAR: Sagaing Division / Alaungdaw Katthapa NP / 22°19.094′N 94°28.823′E / 5.5.2003, ca. 350 m / leg. Boukal et al. (107) ( NHMW) ; 8 males, 7 females: MYANMAR: Sagaing Division / Alaungdaw Katthapa NP / 22°19.094′N 94°28.823′E / 9.- 10.5.2003, ca. 350 m / leg. Boukal et al. (118) ( NHMW, NMPC) ; 1 female: MYANMAR: Sagaing Division / Alaungdaw Katthapa NP / Khaung Din stream / 11.5.2003, ca. 450 m / leg. Boukal et al. (119) // between 22°18.360′N 94°25.937′E / and / 22°19.650′N 94°25.768′E ( NHMW) ; 11 males, 3 females: MYANMAR: Sagaing Division / Alaungdaw Katthapa NP / 22°19.113′N 94°28.518′E / 13.5.2003, ca. 350 m / leg. Schillhammer et al. (122) ( LHCM, NHMW, NMPC, PMCB) ; 3 males, 3 females: MYANMAR (156A) Shan State / on rd Kalaw-Thazi , below (154) / 20°43.796′N 96°29.377′E / ca. 483 m, puddles, 25.06.2004 / leg. Shaverdo & Schillhammer ( NHMW, NMPC) ; 15 males, 9 females: MYANMAR: Shan State (91b) / Shweudaung Wildlife Sanct. / 23°05.129′N 96°13.527′E / 19.7.2002, ca. 360 m / Myint Hlaing & Aung Moe ( NHMW, NMPC) .

Diagnosis. Medium sized, oval oblong species (body length 3.8–4.3 mm). Based on the shape of the median lobe of aedeagus, the new species is probably related to Laccophilus elegans , but differs from that species mainly by the elytral colouration: narrow sinuous transverse subbasal and preapical bands, and large, distinct mediolateral spot. This elytral pattern closely resembles L. hendrichi , but that species has different median lobe with long and narrow apical part.

Description. Colouration: Head and pronotum testaceous. Elytra brownish-black with testaceous pattern consisting of narrow sinuous (especially posteriorly) subbasal transverse band well separated from base of elytra and interrupted before suture, distinct mediolateral spot, sinuous preapical transverse band interrupted before suture, and small apical spot ( Fig. 13 ). Appendages and ventral part testaceous.

Surface structures and sculpture: Head reticulation simple, composed of large irregular polygonal meshes, which are moderately impressed. Frons between eyes with several punctures. Reticulation of pronotum similar to that on head but meshes slightly larger. Pronotum along margins and in posterior angles with numerous setigerous punctures. Reticulation of elytra simple, composed of large irregular polygonal meshes similar to that on pronotum, but slightly more impressed. Meshes complete, with 1–3 clearly-distinct punctures inside of most meshes. Sutural row of punctures reduced to several punctures; discal and lateral rows of punctures present, but especially lateral one barely perceptible. Ventral side reticulated with fine elongate meshes. Prosternal apophysis short and lanceolate. Abdominal ventrites with numerous oblique grooves and several large setigerous punctures, lateral margin bordered. Apical ventrite rounded posteriorly, on disc with several large setigerous punctures and shallow grooves. Male pro- and mesotarsomeres 1–3 weakly dilated with bundle of adhesive setae on ventral side. Claws of pro- and mesotarsi simple and equal.

Male genitalia: Median lobe of aedeagus in lateral view as in Fig. 31 .

Female genitalia: Similar to male; pro- and mesotarsomeres not dilated.

Variability. The specimens of the type series varies slightly in elytral pattern, predominantly in width and sinuosity of transverse bands.

Measurements (N= 40): Body length 3.8–4.3 mm (mean value 4.2 mm, holotype 4.2 mm); width 2.1–2.4 mm (mean value 2.3 mm, holotype 2.4 mm).

Etymology. The new species is dedicated to our colleague Harald Schillhammer (NHMW), a specialist on Staphylinidae , who collected the majority of specimens of this species.

Distribution. Laccophilus schillhammeri is known from several localities in northern Myanmar (Sagaing Region, Shan State) ( Fig. 41 ).

Figures

Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm.

Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm.

Figs. 40–41. Distribution of the species of the Laccophilus javanicus group