Occidenchthonius

Zaragoza, Juan A., 2017, Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae), Zootaxa 4246 (1), pp. 1-221 : 128-129

publication ID

https://doi.org/ 10.5281/zenodo.437611

publication LSID

lsid:zoobank.org:pub:D8594E96-D561-4A37-9073-B138EC7E95A7

DOI

https://doi.org/10.5281/zenodo.5687129

persistent identifier

https://treatment.plazi.org/id/03889E18-FFC0-5596-FF5C-41F9FA1AF928

treatment provided by

Plazi

scientific name

Occidenchthonius
status

gen. nov.

Key to adults of the Occidenchthonius View in CoL species

1 Chelal hand chaetotactic formula 4:4:4, seta ih5 absent. Cheliceral hand lyrifissure ldb present ( O. machadoi View in CoL -group)...... 2

- Chelal hand chaetotactic formula 4:5:4, seta ih5 present. Cheliceral hand lyrifissure ldb present or absent.............. 11

2 Epigean species. Well developed eyes, anterior and posterior eyes with convex lens................................ 3

- Hypogean species. Eyeless or posterior eyes reduced to eye-spots.............................................. 4

3 Fixed chelal finger with 12–16 teeth and movable with 7. Chela stouter: (♂) 4.5, (♀) 3.8–4.7 times longer than deep, length.............................................................................. O. machadoi ( Vachon, 1940) View in CoL

- Fixed chelal finger with 20–21 teeth, movable 9–10. Chela more slender: (♂) 5.1, (♀) 4.6–4.9 times longer than deep.................................................................................... O. canariensis ( Beier, 1965) View in CoL

4 Carapace with a total of 2‒8 preocular and ocular microsetae.................................................. 5

- Carapace with a total of 16 preocular and ocular microsetae............................... O. setosus ( Mahnert, 1993) View in CoL

5 Posterior margin of carapace with 2 (exceptionally 3) setae................................................... 6

- Posterior margin of carapace with 4 (very exceptionally 3 or 5) setae.......................................... 10

6 Eyes absent.............................................................................. O. oromii View in CoL n. sp.

- Eyes present, at least anterior eyes with weak lens, posterior eyes reduced to indistinct spots......................... 7

7 Dorsal face of chelal hand without lyrifissure hd .................................. O. gracilimanus ( Mahnert, 1997) View in CoL

- Dorsal face of chelal hand with lyrifissure hd .............................................................. 8

8 Distal half of movable chelal finger with 9–10 pointed teeth up to slightly proximad of trichobothrium st, basal half with 8–12 rounded and fused vestigial teeth up to level slightly proximad of trichobothria sb ................................. 9

- Distal half of movable chelal finger with 12–15 pointed teeth up to halfway between trichobothria st and sb, basal half with 10–12 rounded and fused vestigial teeth up to halfway between trichobothria sb and b ................. O. mahnerti View in CoL n. sp.

9 Anterior medial margin of carapace prominent; cheliceral spinneret absent in male; stouter and smaller pedipalp: femur (♂) 6.0–6.1, (♀) 5.9 times longer than broad, length (♂) 0.53 mm, (♀) 0.53–0.58 mm; hand (♂) 2.2–2.3, (♀) 2.1 times longer than deep; chela (♂) 5.7–5.8, (♀) 5.0–5.2 times longer than deep, length (♂) 0.71–0.72 mm, (♀) 0.75‒0.76 mm .. O. beieri View in CoL n. sp.

- Anterior medial margin of carapace nearly straight; cheliceral spinneret present in male, hump like; pedipalp femur (♂) 6.8– 7.2, (♀) 6.0–6.9 times longer than broad, length (♂) 0.62–0.65 mm, (♀) 0.63–0.72 mm, hand (♂) 2.4–2.6, (♀) 2.2–2.5 times longer than deep, chela (♂) 5.9–6.5, (♀) 5.5–6.2 times longer than deep, length (♂) 0.79–0.88 mm, (♀) 0.84–0.95 mm .............................................................................. O. tamaran (Mahnert, 2011) View in CoL

10 Smaller, chela length (♀) 0.69 mm, 5.0–5.5 times longer than deep; trichobothrium ist slightly distad to esb .............................................................................................. O. lopezi (Mahnert, 2011) View in CoL

- Larger, chela length (♀) 1.08 mm, 6.1 times longer than deep; trichobothrium ist distinctly distad of esb .............................................................................................. O. tenerifae (Mahnert, 2011) View in CoL

11 Movable cheliceral finger with isolated subapical tooth (di).................................................. 12

- Movable cheliceral finger without isolated subapical tooth (di)............................................... 24

12 Posterior margin of carapace with 4 macrosetae............................................................ 13

- Posterior margin of carapace with 2 macrosetae........................................................... 15

13 Epigean species with normal eyes; carapace with 2 preocular microsetae on each side; fixed chelal finger with 12–16 triangular teeth (not including proximal rounded teeth); smaller: chela length (♂) 0.45–0.57 mm, (♀) 0.50–0.66 mm ............. 14

- Endogean or hypogean species, eyes reduced or absent; carapace with 1 preocular microseta on each side; fixed chelal finger with 17–21 triangular teeth (not including proximal rounded teeth); larger: chela length (♂) 0.89–1.12 mm, (♀) 0.84–1.37 mm ................................................................................ O. cassolai (Beier, 1973) View in CoL

14 Proximal half of fixed chelal finger with 2–5 rounded teeth merging into an evident high bulge of marginal lamina........................................................................................ O. parmensis ( Beier, 1963) View in CoL

- Proximal half of fixed chelal finger with 2–5 rounded and spaced teeth, marginal lamina weakly raised................................................................................................ O. berninii (Callaini, 1983) View in CoL

15 Without eyes or eye-spots............................................................................ 16

- With eyes, anterior pair with convex lens, posterior pair reduced to spots....................................... 23

16 Trichobothrium ist distinctly distad of esb ................................................................ 17

- Trichobothrium ist proximad of esb ....... O. morenoi ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2011) View in CoL

17 Trichobothrium ist proximad of lyrifissure fb ............................................................. 18

- Trichobothrium ist distinctly distad of lyrifissure fb ........................................................ 20

18 Larger and more slender: (♀) pedipalpal femur 7.5 times longer than broad, length 0.88 mm; chela (♀) 7.3 times longer than deep, length 1.23 mm ............................................................ O. sendrai (Zaragoza, 1985) View in CoL

- Smaller and less slender: (♀) pedipalpal femur 5.8–6.5 times longer than broad, length 0.52–0.62 mm; chela (♀) 5.2–5.5 times longer than deep, length 0.70–0.86 mm .................................................................. 19

19 Chelicera with 6 setae and 4–5 lateral microsetae on hand................................. O. thaleri ( Gardini, 2009)

- Chelicera with 6 setae and only 1 lateral microseta on hand............................... O. minutus ( Vachon, 1940) View in CoL

20 Tergite I with 2 setae................. O. ambrosiae ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012) View in CoL

- Tergite I with 4 setae................................................................................ 21

21 Cheliceral hand with 6 setae (excluding microsetae)........................................................ 22

- Cheliceral hand with 7–8 setae (excluding microsetae)........................................................................................... O. bullonorum ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012) View in CoL

22 Larger and slender: pedipalpal femur (♀) 7.5–7.7 times longer than broad, length (♀) 0.93–1.00 mm; chela (♀) 5.9–6.0 times

longer than deep, length (♀) 1.27–1.37 mm ............................................... O. felgueraorum View in CoL n. sp. - Shorter and less slender: pedipalpal femur (♀) 5.5 times longer than broad, length 0.56 mm; chela (♀) 5.2 times longer than deep, length 0.78 mm ................................................. O. villacarrillo ( Zaragoza & Pérez, 2013) View in CoL

23 Lyrifissure ldb present on cheliceral hand, close to seta db. Trichobothrium ist slightly distad of esb and well proximad of lyri- fissure fb ................................................................................. O. anae View in CoL n. sp.

- Lyrifissure ldb absent on cheliceral hand. Trichobothrium ist well distad of esb and slightly proximad of lyrifissure fb ..................................................................................... O. dubius ( Mahnert, 1993) View in CoL

24 Posterior row of carapace with 2 macrosetae.............................................................. 25

- Posterior row of carapace with 4 macrosetae......................................... O. cardosoi (Zaragoza, 2012) View in CoL

25 Chelal hand lyrifissure hp absent....................................................................... 26

- Chelal hand lyrifissure hp present...................................................................... 27

26 Carapace without preocular microsetae.................................................... ( O. verai View in CoL -group) 28

- Carapace with 2 preocular microsetae on each side....................................... O. pinai (Zaragoza, 1985) View in CoL

27 Smaller, epigean, species: pedipalp femur (♀) 5.1–5.3 times longer than broad, length 0.44–0.47 mm; chela (♀) 3.7–3.9 times longer than deep, length 0.61 mm .......................................................... O. serranoi View in CoL n. sp.

- Larger, hypogean, species: pedipalp femur (♀) 8.7–8.8 times longer than broad, length 1.28–1.30; chela (♀) 7.5–7.9 times lon- ger than deep, length 1.72–1.82............................................................ O. ortunoi View in CoL n. sp.

28 Subterranean species without eyes...................................................................... 29

- Epigean species with eyes, anterior pair with convex lens........................................ O. gardinii View in CoL n. sp.

29 Chelicera with 6 macrosetae and 1–2 (rarely 3) lateral microsetae. Trichobothrium ist distinctly distad of esb ........... 30

- Chelicera with 6 macrosetae and 4–5 lateral microsetae. Trichobothrium ist level with esb ................................................................................................ O. giennensis ( Zaragoza & Pérez, 2013) View in CoL

30 Smaller. Chela length about 0.70 mm or less.............................................................. 31

- Larger. Chela length about 1.00 mm or more.............................................................. 32

31 Fixed chelal finger with 14–15 teeth; stouter and shorter pedipalp: femur (♂) length 0.39–0.42 mm, 4.8–5.0 times longer than broad, chela (♂) length 0.58–0.60 mm, 4.8–5.0 times longer than deep........................... O. montagudi View in CoL n. sp.

- Fixed chelal finger with 19 teeth; more slender and longer pedipalp: femur (♂) length 0.54 mm, 6.0 times longer than broad, chela (♂) length 0.71 mm, 5.7 times longer than deep.......................................... O. lencinai View in CoL n. sp.

32 Trichobothrium ist proximad of or level with lyrifissure fb; smaller species, pedipalpal femur (♀) 5.7–7.5 times longer than broad, length 0.66–0.94 mm, chela (♀) length 0.94–1.29 mm ................................................ 33

- Trichobothrium ist distinctly distad of lyrifissure fb; larger species, pedipalpal femur (♀) 7.9–8.6 times longer than broad, length about 1.10 mm, chela (♀) length 1.40–1.53 mm ......................................................41

33 Movable chelal finger with 3–8 rounded vestigial teeth without dental canals in basal half......................... 34

- Movable chelal finger with 9–12 rounded vestigial teeth without dental canals in basal half........................ 37

34 Trichobothrium ist distinctly proximad of lyrifissure fb ..................................................... 35

- Trichobothrium ist level with lyrifissure fb ..................................................... O. riopar View in CoL n. sp.

35 Lyrifissure ldb present on cheliceral hand. O. cazorlensis Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001 View in CoL

- Lyrifissure ldb absent on cheliceral hand................................................................. 36

36 More slender species, pedipalpal femur (♀) 6.9–7.1 times longer than broad......................... O. mateui View in CoL n. sp.

- Stouter species, pedipalpal femur (♀) 6.1 times longer than broad................................ O. hoerwegi View in CoL n. sp.

37 Cheliceral fixed finger with 2 large distal teeth............................................................ 38

- Cheliceral fixed finger with only 1 large distal tooth........................................................................................... O. torremarinae ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012) View in CoL

38 Smaller: (♀) pedipalpal femur length 0.66–0.75 mm, 5.7–6.4 times longer than broad; chela (♀) length 0.94–1.10 mm, 5.2–5.5 times longer than deep.................... O. perezi ( Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2011) View in CoL

- Larger: (♀) pedipalpal femur length 0.80–0.99 mm, 6.4–7.5 times longer than broad; chela (♀) length 1.06–1.38 mm, 5.7–6.5 times longer than deep............................................................................... 39

39 Male spinneret prominent........................................................... O. verai (Zaragoza, 1985) View in CoL

- Male spinneret absent or strongly reduced................................................................ 40

40 Lyrifissure ldb of cheliceral hand present.............................................. O. ventalloi ( Beier, 1939) View in CoL

- Lyrifissure ldb of cheliceral hand absent.................................... O. espanyoli ( Zaragoza & Pérez, 2013) View in CoL

41 Movable chelal finger with 14 vestigial teeth without dental canals in basal half....................... O. murcia View in CoL n. sp.

- Movable chelal finger with only 6–8 vestigial teeth without dental canals in basal half............................ 42

42 Lyrifissure ldb of cheliceral hand present................................................................................................... O. ruizporteroae (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) View in CoL .

- Lyrifissure ldb of cheliceral hand absent......................................................... O. ebo View in CoL n. sp.

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