Amphidraus zaque, Galvis, William, 2017

Galvis, William, 2017, Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation, Zootaxa 4286 (1), pp. 1-40 : 25-26

Amphidraus zaque sp. nov.

Figs 14a–d , 19a–b , 23d–e , 25e, 25k , 27

Types. Holotype: male from Finca Alirio González , Vereda Montealegre, Topaipí, Cundinamarca, Colombia, 1377 m, 5.400916°N, 74.294016°W, 18–23.X.2012, M. Medrano, A. García ( ICN –Ar 3378) . Paratype: one female from Finca Pedro Méndez , Vereda Montealegre, Topaipí, Cundinamarca, Colombia, 1511 m, 5.399216°, 74.299716°W, M. Medrano, A. García ( ICN –Ar 3379).

Etymology. The epithet is a noun in apposition that honors one of the four supreme rulers of the Muisca Confederation, ancient inhabitants of the central Andean highlands of the present-day Colombia’s Eastern Range: the Zaque , the Zipa , the Iraca and the Tundama .

Diagnosis. Males of A. zaque sp. nov. are similar to those of A. semicanus ( Simon, 1902) , A. sikuani sp. nov., A. sotairensis sp. nov. and A. tanimuca sp. nov. by their bilobed RTA. However, they can be distinguished from those by their halberd-like RTA with a long embolic filament (EF), and a bump in the trochanter ( Figs 14a–b ). Females of A. zaque sp. nov. can be easily distinguished from those of the remaining known females of the genus for the longer and coiled copulatory ducts, with more than three coils before the insertion in the spermathecae, with the first part of the copulatory ducts upright, and slender spermathecae with a long anterior lobe after the fertilization ducts ( Figs 14c–d , 25e, 25k ).

Description. Male (holotype). Total length: 2.79. Carapace dark brown with a dorsal oval yellowish mark, 1.53 long, 1.14 wide, 0.84 high ( Fig. 19a ). OC black, 0.63 long. Anterior eye row 1.17 wide and posterior 1.05 wide. Sternum dark brown, 0.57 long, 0.49 wide. Labium dark brown, 0.18 long, 0.24 wide. Chelicerae reddish brown with four retromarginal and two promarginal teeth. Palp brown with distal femur and patella with long white hairs, with a halberd-like RTA, and long tegular lobe (TL), long embolic filament (EF), and a bump in the trochanter (black arrow in the Fig. 14a ) ( Figs 14a–b , 23d–e ). Legs 4132, all yellow with dark brown marks. Leg macrosetae: femur, II d 1 di, p 1 di; III d 2 di, p 1 di; patella, III–IV r 1 me; tibia, I v 2-2 -2; II v 1-2 -1, p 1 di; III–IV v 0-1-1, p 0-1-1, 0-1-1; metatarsus, I v 2-2; II v 2-2, p 1 di; III v 1 -0-1, p 1-0-2, r 1-0-2; IV v 1 di, p 1-0-2, r 1-0-2. Abdomen brown with a reticulated light brown pattern of marks ( Fig. 19a ).

Female (paratype). Total length 3.08. Carapace yellow with two posterior dark brown longitudinal stripes, 1.47 long, 1.07 wide, 0.82 high ( Fig. 19b ). OC black, 0.65 long. Anterior eye row 1.12 wide and posterior 0.99 wide. Sternum dark yellow, 0.59 long, 0.44 wide. Labium dark yellow, 0.15 long, 0.19 wide. Chelicerae dark yellow, with five tiny retromarginal and two promarginal teeth. Legs 4312, all yellow. Leg macrosetae: femur, II–III d 1 di; patellae, III–IV r 1 me; tibiae, I v 2-2 -2; II v 2-2 -2, p 1 di; III v 1 pr, p 1-1, r 1-1; IV v 1 -0-1, p 1-1, r 1- 1; metatarsus, I v 2-2; II v 2-2, p 1 di; III v 1 -0-1, p 1-0-2, r 1-0-2; IV p 1-0-2, r 1-0-2. Abdomen gray, with a dark brown reticulated pattern of marks ( Fig. 19b ). Epigyne ( Figs 14c–d , 25e, 25k ) with anterior copulatory openings, a posterior single medial pocket, longer and coiled copulatory ducts, with more than three laps until the insertion in the spermathecae, with the first part of the copulatory ducts upright, and medial to anterior slender spermathecae with a long anterior lobe after the fertilization ducts.

Comments. The type material was collected in a pitfall trap in a corn crop.

Distribution. Colombia (Cundinamarca) ( Fig. 27 ). Known altitudinal distribution: 1377–1511 m.

Figures

FIGURES 14 a – d. A. zaque sp. nov., a) left male palp, ventral view; b) same, retrolateral view (black arrow showing the bump on the trochanter); c) epigyne, ventral view; d) same, cleared, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.30 mm.

FIGURES 19 a – d. A. zaque sp. nov., a) male holotype (ICN – Ar 3378); b) female paratype (ICN – Ar 3379). A. zipa sp. nov., c) male holotype (ICN – Ar 3363); d) female paratype (ICN – Ar 3364). Scale bars: 0.50 mm.

FIGURES 23 a – g. Amphidraus spp, left male palp. A. tundama sp. nov., a) ventral view; b) same, retrolateral view; c) palpal femur, showing the proventral bump (PvFB, black arrow), ventral view. A. zaque sp. nov., d) ventral view; e) same, retrolateral view. A. zipa sp. nov., f) ventral view; g) same, showing the proventral femoral widening (PvFw, black arrow) in the femur, retrolateral view.

FIGURES 25 a – l. Amphidraus spp, epigyne. A. sikuani sp. nov., a) ventral view; g) disposition of the internal structures. A. somondoco sp. nov., b) ventral view; h) disposition of the internal structures. A. sua sp. nov., c) ventral view; i) disposition of the internal structures. A. tundama sp. nov., d) ventral view; j) disposition of the internal structures. A. zaque sp. nov., e) ventral view; k) disposition of the internal structures. A. zipa sp. nov., f) ventral view; l) disposition of the internal structures.

FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments.