Tokyosoma breviprocessum, Mikhaljova, Elena V., Golovatch, Sergei I. & Chang, Hseuh-Wen, 2010

Mikhaljova, Elena V., Golovatch, Sergei I. & Chang, Hseuh-Wen, 2010, The millipede family Diplomaragnidae in Taiwan, with descriptions of nine new species (Diplopoda, Chordeumatida), Zootaxa 2615, pp. 23-46 : 32-34

Tokyosoma breviprocessum sp. nov.

Figs 20–22 .

Material examined: Holotype: 1 male ( NMNS –6345–010, D–0617), Taiwan, Nantou County, Huisun timberland, 24 April 1998, leg. S.H. Wu. Paratypes: 1 male ( IBSS), D–0646, Taiwan, Nantou County, Huisun timberland, December 1997, leg. S.H. Wu. 1 male ( NMNS –6345–011, D–0219), 1 male (NSYSUB), D–0225, Taiwan, Nantou County, Huisun timberland, 31 January 2000, leg. S.H. Wu.

Diagnosis: Differs from congeners mainly by the shape of the posterior gonopod lateral coxal branch carrying two apical horn-shaped processes and one frontal finger-shaped process in the middle portion, by the form of the posterior gonopod colpocoxite with a distal, short, pointed outgrowth, as well in a half-gobletshaped coxal process of male leg 10.

Description: Male. Length about 12 mm, width with paraterga 1.0 mm. Coloration in alcohol beige throughout, ocellaria brown (possibly faded).

Body with 32 segments. Head covered with sparse, relatively long and short setae. Each eye patch composed of at least 27 ocelli. Collum semi-circular. Body width gradually increasing until somite 7, body parallel-sided on somites 8–15, thereafter gradually tapering. Beginning from somite 4, rounded paraterga normally developed, growing increasingly less distinct towards hind part of body, strongly reduced on somites 27 and 28, absent from somites 29–31. Paraterga 2 and 3 small. Metazonital macrochaetae in a transverse row on somites 30 and 31, like an elongate (to different degrees) triangle on preceding somites. Macrochaetae in anterior part of body relatively long, pointed apically, but in middle and posterior parts of body anterolateral macrochaetae short (baton-shaped), posterolateral ones long, medial macrochaetae somewhat shorter than posterolateral ones, all blunt. Metazonites with two very low knobs placed near axial suture on each side and with an oblong bolster on paraterga.

Leg pairs 3–7 somewhat enlarged. Leg pairs 6 and 7 with a group of funnel-shaped tarsal papillae apically near claw. Claw of leg pairs 6 and 7 at base with a long setoid filament ventrally, but without additional dorsal claw. Postgonopodal legs (including leg pairs 10 and 11) without tarsal papillae. Claw of legs 10 and 11 at base with one small additional claw dorsally and a long setoid filament ventrally. Claw of midbody and hindmost legs at base with one small additional claw dorsally and a long setoid filament ventrally.

Legs 10 and 11 with coxal glands. Coxa 10 papillate caudally, caudoventrally with a half-goblet-shaped process ( Fig. 20 ). Trochanter 10 with a small ventral outgrowth setose apically; right outgrowth somewhat smaller than left one. Coxa 11 without processes. Trochanter 11 with a caudal finger-shaped process papillate mesally and rounded apically.

Anterior gonopod telopodite 1-segmented, flagelliform, densely beset with cuticular spinules, its distal part positioned inside sheaths with very strongly elevated and tightly closed edges (e) ( Fig. 21 ). These sheath edges like high plates in lateral view. Telopodite base and distal part of coxosternum tightly attached to adjacent mesal portion of posterior gonopod. Posterior gonopod colpocoxites fused sub-basally, curved caudad. Colpocoxites distally with short pointed processes (c), laterally with thin blades (b). Mesal sheath processes of posterior gonopod colpocoxites fused medially into a single cup-shaped structure (ms) covered with long pointed spinules. Lateral sheath processes of colpocoxites (lp) cup-shaped, carrying pointed spinules caudally.

Posterior gonopod angiocoxite with a globule, but without process in posterior view. Posterior gonopod coxal part ( Figs 21-22 ) with a long lateral branch (lb) curved caudad and carrying two apical horn-shaped processes, one mesocaudal (mbp), other lateral (lbp), as well as one frontal finger-shaped process (f) in middle portion. Basal part of this branch fused with both colpocoxite and anterior angiocoxite.

Angiocoxite depressed centrally in anterior view ( Fig. 22 ), supplied with a long process (ap); distal portion of this process penetrating colpocoxite and visible outside in lateral view. Posterior gonopod telopodite 2-segmented; trochanteroprefemur hollow for accommodation of lateral branch (lb) mesocaudally; femur like a knob (part of femur visible through wall of trochanteroprefemur of posterior gonopod telopodite).

Female unknown.

Name: The specific epithet refers to the short distal outgrowth of the posterior gonopod colpocoxite.

Figures

FIGURES 20 – 22. Tokyosoma breviprocessum sp. nov., male paratype. 20, coxa and trochanter 10, front view; 21, gonopods, caudal view; 22, gonopods, front view; e, edge of posterior gonopod colpocoxite sheath; c, distal process of posterior gonopod colpocoxite; b, posterior gonopod colpocoxite lateral blade; ms, mesal sheath processes of posterior gonopod colpocoxites; lp, lateral sheath process of posterior gonopod colpocoxite; lb, posterior gonopod lateral coxal branch; mbp, mesocaudal apical process of the posterior gonopod lateral coxal branch; lbp, lateral apical process of the posterior gonopod lateral coxal branch; f, finger-shaped process of the posterior gonopod lateral coxal branch; ap, anterior process of posterior gonopod angiocoxite. Scale in mm.