Lymantes fowleri Anderson, 2016

Anderson, Robert S., 2016, A Taxonomic Revision of the GenusLymantesSchönherr, 1838 (Coleoptera: Curculionidae: Molytinae: Lymantini) in the United States Ofamerica, The Coleopterists Bulletin 70 (1), pp. 111-124 : 111-124

Lymantes fowleri Anderson , new species ( Figs. 10–11 , 17 , 22 )

Diagnosis. Eyes present as small curved row of 2–3 individually indistinct facets on slight ocular swelling ( Fig. 17 ). Punctures of body generally moderately large, widely spaced, those of pronotal disc moderately deep, moderately large, widely spaced, the distance between punctures at least one-half diameter of a puncture or more ( Fig. 11 ). Punctures of striae of elytral disc moderately large, deep and linearly arranged, interstriae distinct, about as wide as a strial puncture ( Fig. 11 ). Dorsal vestiture indistinct, minute and very fine, each seta much shorter than diameter of elytral strial puncture ( Fig. 10 ). Legs moderately long, not spindly, hind femora about 5X as long as wide, apex reaching to suture between ventrites 2 and 3. Aedeagus as in Fig. 22 .

Description. Male, length 2.5–3.0 mm, width 1.1–1.3 mm. Color medium orange to red brown throughout. Head globose, minutely finely reticulate. Eyes present as small curved row of 2–3 individually indistinct facets on slight ocular swelling. Rostrum with antennae inserted slightly beyond apical 1/3 of length, slightly wider in dorsal view beyond antennae insertion, scrobes narrowly visible at point of antennal insertion in dorsal view. Rostrum about 0.7X length of pronotum, slightly arcuate in lateral view, dorsally moderately deeply, moderately densely linearly punctate, punctures separated by less than their own diameter, row of punctures above scrobe coalescent, forming shallow groove. Ventrally irregularly punctate, most punctures coalescent, with median posterior extension represented by narrowly separated lateral cariniform ridges, extended to base of rostrum. Antennae with scape almost reaching base of rostrum, funicle with articles 1 and 2 elongate, subequal in length, articles 3–6 subquadrate, article 7 wider than longand wider towards apex. Pronotum with width about 0.8X length, lateral margins slightly arcuate, widest at about midlength, subapical constriction not evident dorsally, distinctly impressed laterally, disc with punctures moderately deep, moderately large, widely spaced, the distance between punctures at least one-half diameter of a puncture or more, each puncture with minute, fine erect seta. Elytra about 1.8X as long as wide, lateral margins subparallel from behind humeri to apical 1/4, humeri not pronounced, disc with striae with large, deep and linearly arranged punctures numbering 15–20 per complete discal stria, interstriae very finely punctate, shining, narrower than width of a strial puncture, punctures each with fine, minute erect seta, each seta much shorter than diameter of elytral strial puncture. Striae 1–6 complete to base, 7 and 8 fused opposite metacoxae, not continuous to base, stria 10 terminated opposite metacoxa. Venter deeply, moderately densely, regularly punctate, punctures well-spaced, except punctures lacking on ventrites 3 and 4 and slightly larger on ventrite 5. Abdominal ventrites 1 and 2 very long, 1 longer than ventrites 3–5 combined, 2 about as long as ventrites 3 and 4 combined, ventrites 3 and 4 very short, subequal in length, ventrite 5 about 1.5X as long as 3 and 4 combined. Legs long and slender, sparsely shallowly punctate, punctures well-spaced,, femora extended to suture between ventrites 2 and 3, about 4X as long as wide, tibiae straight, slender, about ¾ as long at femora. Tarsi slender, ventrally with scattered setae, tarsal claws very small, fine. Aedeagus with pedon short, very slightly longer than wide; with internal sac with extensive internal sclerotization in form of a pair of distal arciform sclerites, basally with “U-shaped” band-like sclerite basal to an irregularly shaped median complex; struts short, about 1.5X length of pedon.

Female differs from male as follows: length 2.5–3.2 mm, width 1.1–1.3 mm. Rostrum with antennae inserted slightly proximad of apical 1/3 of length. Female not dissected.

Derivation of Species Name. I take pleasure in naming this species after country singer Kevin Fowler, of Austin, Texas, the best non-mainstream country music artist out there. Kevin is a resident of the Austin area, an avid outdoorsman, and proud Texan.

Natural History. Specimens were collected in various kinds of leaf litter, often in riparian situations or in moist canyons throughout the Edwards Plateau. Specimens have been collected from November through May.

Distribution. This species is known from central Texas.

Material Examined. Holotype male ( CMNC), Texas: Gonzales County, Palmetto State Park , east entry to park, 17–18 Apr 1989, R. Anderson, oak- Celtis leaf litter . Paratypes as follows. Same data as holotype (11m, 9f, ASUHIC, BMNH, CMNC, CWOB, TMMC, USNM) . Bandera County, Lost Maples State Natural Area , 26 Apr 1986, P.W. Kovarik, berlese leaf litter (1 female, TAMU) . Bandera County, Lost Maples State Natural Area , 28 Apr- 2 May 1987, R.Anderson, berlese leaf/log litter (4 males, 1 female, CMNC, SEMC) . Bandera County, Lost Maples State Natural Area, Maple Trail , 9–10 Mar 2001, C.W. O’ Brien, sifted maple litter (7 males, 5 females, CWOB) . Bandera County, Lost Maples State Natural Area , 27 Mar 1999, E.G. Riley, berlese big toothed maple litter (1 female, EGRC) . Bandera County, Bandera Homestead , 5 Apr 1990, W.F. Barr, oak litter in limestone outcropping (1 male, WFBM) . Bell County, C.B. Cave, Fort Hood , 21 Apr 1998, J. Reddell, M. Reyes (1 female, TMMC) . Bell County, Camp Kachina , 4 mi. W. Temple, 5 Mar 1995, E.G. Riley, berlese (1 female, EGRC) . Brazos County, Koppe’ s Bridge ca. 5 mi. S.W. College Station, 11 Feb 1989, R.Anderson, berlese riparian litter (3 males, 4 females, CMNC) . Brazos County, Koppe’ s Bridge ca. 5 mi. S.W. College Station, 21–22 Jan 1989, R.Anderson, berlese riparian litter (1 male, 1 female, CMNC) . Brazos County, Koppe’ s Bridge ca. 5 mi. S.W. College Station, 28 Feb 1988, R.Anderson, berlese riparian litter (1 male, CMNC) . Brazos County, Koppe’ s Bridge ca. 5 mi. S.W. College Station, 6–12 Apr 1987, R.Anderson, berlese riparian litter (3 males, 5 females, CMNC, CWOB) . Brazos County, Koppe’ s Bridge ca. 5 mi. S.W. College Station, 6 Mar 1988, R.Anderson, berlese riparian litter (3 males, 2 females, CMNC) . Brazos County, Koppe’ s Bridge ca. 5 mi. S.W. College Station, 15 Jan 1989, R.Anderson, berlese riparian litter (2 males, CMNC) . Brazos County, Koppe’ s Bridge ca. 5 mi. S.W. College Station, 21 Mar 1988, R.Anderson, berlese riparian litter (1 male, CMNC) . Brazos County, Koppe’ s Bridge , 5 mi. S.W. College Station, 5 Feb 1994, E.G. Riley, berlese forest litter (1 female, CWOB) . Brazos County, Koppe’ s Bridge , 5 mi. S.W. College Station, 20 Nov 1993, E.G. Riley (2 males, 2 females, CWOB, EGRC) . Brazos County, Koppe’ s Bridge , 5 mi. S.W. College Station, 5 Feb 1994, E.G. Riley (1 female, EGRC) . Brazos County, Koppe’ s Bridge , 5 mi. S.W. College Station, 26 Mar 1994, E.G. Riley, berlese forest litter (2 females, EGRC) . Brazos County, Koppe’ s Bridge , 5 mi. S.W. College Station, 15 Jan 1994, E.G. Riley (1 female, EGRC) . Brazos County, Koppe’ s Bridge , 5 mi. S.W. College Station, 4 Feb 1995, E.G. Riley (2 females, EGRC) . Burleson County, Junction Hwy 50 and Yegua Creek near Clay , 18 Feb 1995, E.G. Riley, berlese river drift (1 female, EGRC) . Colorado County, Columbus , 17 May 1989, R.Anderson, berlese riparian litter (2 females, CMNC) . Colorado County, Columbus , 22 Mar 1988, R.Anderson, berlese riparian litter (1 male, CMNC) . Fayette County, 0.5 mi. W. La Grange , 22 Mar 1988, R.Anderson, berlese riparian litter (1 female, CMNC) . Travis County, vic. Long Hollow Creek , 30°27′30″N 97°52′30″W, 7 May 1994, E.G. Riley, berlese forest litter (2 males, 5 females, TAMU) . Travis County, Tooth Cave , 18 Mar 2009, P. Paquin, K. O’ Connor, surface sifting (7 males, 4 females, ASUHIC, CMNC, TMMC) . Uvalde County, Concan, Neal’ s Lodge Area , 9 Apr 1995, E.G. Riley, berlese forest litter (6 males, 1 female, CWOB, EGRC) . Washington County, Washingtonon-the-Brazos State Park , 6 Apr 1988, R.Anderson, berlese riparian litter (1 female, CMNC) .

Figures

Figs. 4–13. Lymantes species, lateral and dorsal habitus. 4–5) L. scrobicollis; 6–7) L. repens; 8–9) L. nadineae;

Figs. 14–18. Lymantes species, head, lateral view. 14) L. scrobicollis; 15) L. repens; 16) L. nadineae; 17) L. fowleri; 18) L. obrieni.

Figs. 19–23. Lymantes species, male aedeagus, dorsal view. 19) L. scrobicollis; 20) L. repens; 21) L. nadineae;