Dichotomius (Luederwaldtinia) henripittieri,

Montoya-Molina, Santiago & Vaz-De, Fernando Z., 2021, Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae), European Journal of Taxonomy 734, pp. 1-64: 31-33

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Dichotomius (Luederwaldtinia) henripittieri

sp. nov.

Dichotomius (Luederwaldtinia) henripittieri  sp. nov.


Fig. 10View Fig, Map 11


Dichotomius henripittieri  sp. nov. differs from other species of the group by the following combination of characters: clypeal teeth short and rounded. Clypeo-genal junction not emarginate. Cephalic carina, wide as high, median tubercle rounded at apex. Dense, ocellate punctures covering interocular surface. Pronotum twice as wide as long. Anterior medial disk with weak declivity, not truncate. Impressed fine punctures restricted to antero-lateral edges. Row of uninterrupted ocellate punctures along pronotal margin. Interstriae microsculpture shagreened.


Name based on the locality where this new species was collected, Henri Pittier National Park, Venezuela.

Material examined

Holotype VENEZUELA • ♂; Aragua, Mario Briceño Iragorry , PN. Henri Pittier , Est. Bio. Periquito ; 1145 m a.s.l.; 10º20′51″ N, 67º41′17″ W; human faeces; 18–20 Aug. 2010; R. Acconcia and J. Gámez leg. [white, p]; CEMT.GoogleMaps 

Paratypes VENEZUELA • 3 ♂♂, 3 ♀♀; same collection data as for holotpye [white, p]; CEMT  .



MEASUREMENTS. Length: 19.36 mm.

HEAD. Wider than long. Anterior edge surface weakly rugose, extending posteriorly not reaching base of cephalic carina. Clypeo-genal surface covered by ocellate punctures spaced by one time diameter of a puncture. Interocular surface with ocellate punctures next to eyes ( Fig. 10A, CView Fig). Clypeus rounded, weak central emargination separating two rounded clypeal teeth. Ventral clypeal process longitudinally enlarged, cylindrical (see Fig. 2DView Fig). Cephalic carina as wide as high, posterior surface shagreened microsculpture with ocellate shallow punctation. Median tubercle three times as high as lateral tubercles and curved posteriorly, apex rounded never bifurcate. Lateral tubercles obtuse at apex, vertically flattened ( Fig. 10CView Fig).

PRONOTUM. Twice as wide as long, and as wide as elytra. Anterior medial disc with weak declivity, not truncate. Median disc shiny and smooth, lacking deep punctures and having at most fine punctures. Antero-lateral surface with dense well-impressed simple punctures, scattered ocellate punctures also visible. Row of ocellate punctures along margins, punctures denser at anterior and posterior margins edges. Hypomeron medial disc punctation separated by three times width of a puncture, never smooth or impunctate. External margin bearing single row of organized long setae (see Fig. 2FView Fig).

ABDOMEN. Prosternum and mesosternum with ocellate setigerous punctures. Metepisternum surface shagreened, displaying ocellate setigerous punctures, pilosity sparce. Metasternum setigerous punctures restricted mainly to lateral and anterior areas of mestasternal lobe, puncture diameter variable, medial disc smooth. Margins of mesocoxa surrounded by multiple ocellate punctures, reaching lateral portions of metasternum ( Fig. 10DView Fig). Lateral surface of metasternum, with ocellate setigerous punctures separated by three times width of a puncture. Ocellate punctures along sternites anterior margins. Pygidium wider than long, basal margin rugose, bearing ocellate punctures in basal half.

ELYTRA. Interstriae microsculpture shagreened, with smooth fine scattered punctures. Striae wellimpressed with ocellate punctures separated by less the diameter of a puncture on elytral disc ( Fig. 10AView Fig).

LEGS. Metacoxa posterior edge bearing single row of disorganized ocellate setigerous punctures, separated by twice diameter of a puncture. Profemora displaying aggregation of setigerous punctures on distal ¾.


Morphological variation

Females ( Fig. 10BView Fig)

As males except for the following characters:

HEAD. Completely transversely rugose, denser on anterior portion. Clypeus with strong central emargination separating two acute clypeal teeth. Cephalic carina bituberculate, tubercles narrow, almost fading ( Fig. 10BView Fig).

PRONOTUM. Surface covered by fine punctures, denser on antero-lateral areas. Punctuation on margins ocellate and dense, discernible. Hypomeron setae longer and denser than in males.


This species was collected in the Henri Pittier National Park located on the ridge of the Cordillera de la Costa of Venezuela. The vegetation of the area is characterized by extensive cloud forest.