Dichotomius (Luederwaldtinia) fornicatus ( Luederwaldt, 1931 ),

Montoya-Molina, Santiago & Vaz-De, Fernando Z., 2021, Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae), European Journal of Taxonomy 734, pp. 1-64: 27-29

publication ID

https://doi.org/10.5852/ejt.2021.734.1233

publication LSID

lsid:zoobank.org:pub:663A0C2D-E3CF-4B0C-8BA1-566B174829A2

DOI

http://doi.org/10.5281/zenodo.4545851

persistent identifier

http://treatment.plazi.org/id/03850669-FF8A-FFD2-FE33-20A03932CCBE

treatment provided by

Plazi

scientific name

Dichotomius (Luederwaldtinia) fornicatus ( Luederwaldt, 1931 )
status

 

Dichotomius (Luederwaldtinia) fornicatus ( Luederwaldt, 1931)  Fig. 9View Fig, Map 6

Pinotus opacipennis var. fornicatus Luederwaldt, 1931: 302  (original description).

Dichotomius bosqi Pereira, 1941: 263  (original description). Syn. nov.

Dichotomius opacipennis var. fornicatus  – Blackwelder 1944: 207.

Dichotomius bosqi Martínez 1959: 89  .

Diagnosis

Dichotomius fornicatus  is separated from other species in the group by the following combination of characters: small body size, less than 15 mm. Dorsal and ventral surface dark blue to black. Head cephalic carina, distinctly arcuate in dorsal view, middle edge slightly forward of lateral. Pronotum with abrupt anterior declivity when viewed laterally, middle surface deeper, apex of the declivity with two well-developed lobes. Elytra opaque, shagreened microsculpture.

Material examined

Holotype of P. fornicatus  ARGENTINA • ♀; “Republ. Argentine CHACO DEL ESTERO, RIO SALADO” [white with black superior and inferior margins, p]/ “DÉCEMBRE” [white, p]/ “MÚSEUM PARIS 1938 COLL. A. BOUCOMONT” [green with black margins, p]/ “ Pinotus opacipennis Lueder.  Forma a. Luederw. Det. 1931” [white, hw]; MNHN. 

Holotype of D. bosqi 

ARGENTINA • ♂; “ Sgo del ESTERO . Rio Salado. WAGNER col.” [white with black margins, p]/ “TIPO” [red with black margins, p]/ “ Pinotus bosqi P. Pereira  det. 940” [white with black margins, phw]/ “23.030” [white, hw]; MZSP. 

Paratype

ARGENTINA • 1 ♀; “ Sgo del ESTERO . Rio Salado. WAGNER col.” [white with black margins, p]/ “ ♀” [white, hw]/ “Paratypo” [red, hw]/ “ Pinotus bosqi ♀ P. Pereira  det. 941” [white with black margins, phw] /“289” [red rounded, hw]; MLP  .

Additional material

ARGENTINA – Cordoba • 1 ♀; La Para. C/esc; Dec. 1996; Monteresino leg.; CEMT  1 ♂; C. a mano ; Nov. 1990; Monteresino; CEMT  4 ♀♀; Salta, Salado River ; J.V. González and S. Macapilo leg.; 18 Jan. 2010; Snizek leg.; CEMT  1 ♂; Rosa. Front. El Naranjo ; A. Martínez leg.; MACN  1 ♂; Tablillas ; Dec. 1932 – Sep. 1933; CEMT. – Santiago del Estero  2 ♀♀; Campo Gallo, S. del. Estero ; Mar. 1943; A.F.P. B.M; BMNH 1948-159  2 ♂♂, 1 ♀; Choya ; Oct. 1961; MLP  1 ♂, 4 ♀♀; El Charco ; Feb. 1956; MLP  1 ♂; Matará ; 12 Dec. 1939; Biraben-Bezzi leg.; MLP  2 ♂♂; Rio Salado ; Wagner leg.; MLP  2 ♀♀; Thermas de Rio Hondo ; 27–28 Nov. 1979; C. and M. Vardy B.M. leg.; BMNH 1980-67  1 ♀; Tucumán; Jan. 1947; CEMT  1 ♀; Dec. 1948; CEMT  1 ♀; Campo Bello ; Mar. 1943; MACN  1 ♂, 1 ♀; San Pedro Colalao ; Feb. 1949; CEMT  1 Ƌ; “ METATIPO”/ “Campo Gallo, S. del Estero III-1943 A.F.P.”/ “ Pinotus  Ƌ bosqi P. Pereira  det. 948/B.M. 1948-159.” / “NHMUK 010847032”; BMNH  1 ♀; “ METATIPO” /“Campo Gallo, S. del Estero III-1943, A.F.P.” /“ Pinotus  bosqi P. Pereira  det. 948 /B.M. 1948-159” / “NHMUK 010847033”; BMNH  .

BOLIVIA • 1 ♀; Cordillera, Tunalito ; 8 Nov. 1946; Maldonado leg. and coll. Martinez.; CEMT. – Santa Cruz  1 ♂, 1 ♀; Ascención ; Nov. 1963; MLP  1 ♀; Ñuflo de Cháves ; Nov. 1963; MLP  1 ♂; Yoay ; Dec. 1933; G.L. Harlington leg.; CEMT  1 ♂; Tarija, Res.Priv. Corbalán ; 298 m a.s.l.; 21º39′ S 62º28′ W; chaco seco. trap. caída cebo humano; 12 Dec. 2004; C. Hamel and M. Aliaga leg.; CEMTGoogleMaps  2 ♂♂, 2 ♀♀; between Caiza and Creveaux; 536 m a.s.l.; 21º48′53″ S, 63º26′53″ W; blue light; OUMNH-2005-065; 05-I-2005; Mann, Hamel and Herzog leg.; CEMTGoogleMaps  1 ♂; Yoay and Mandipecua [Yoay y Mandeyapecua]; collector 479; Nov. 1932; Press by WCH rington; CEMT  .

PARAGUAY – Boquerón • 2 ♂♂, 3 ♀♀; Loma Plata , Chaco ; 8–29 Jan. 1995; CEMT  . – Caaguazú • 1 ♀; Capiata ; 20 Nov. 2001; C. Aguilar leg.; CEMT  .

COUNTRY UNKNOWN • 1 ♂; no data; CEMT  1 ♂, 1 ♀; B.M.; BMNH 1948-159  .

Redescription

Male

MEASUREMENTS. Length: 13.35 mm. Dorsal and ventral surfaces dark blue to black. Pronotum and ventral surfaces shiny, elytra opaque.

HEAD. Surface rugosity not extending posteriorly to base of cephalic carina; interocular surface with well-impressed punctures near eyes. Cephalic carina, wider than high, distinctly arcuate in dorsal view ( Fig. 9GView Fig), middle edge slightly forward of lateral. Median tubercle approximately twice as high as lateral tubercles, conic in shape and slightly curved posteriorly. Ventral clypeal process cylindrical (see Fig. 2DView Fig).

PRONOTUM. Wider than long. Antero-median declivity of pronotal disc with two underdeveloped lobes at sides when viewed laterally ( Fig. 9AView Fig). Pronotal disc shiny with dense deep punctures, separated by diameter of a puncture; margins rugose, denser at antero-lateral edges. Hypomeron medial disc smooth and shiny, external margin with single row of organized and scattered setae of same length.

ABDOMEN. Mesosternum covered by large ocellate punctures separated by diameter of a puncture. Metasternum antero-medial edge smooth and shiny. Ocellate punctures restricted mainly to lateral edges, punctures separated by width of a puncture. Small ocellate punctures surrounding margin of mesocoxa ( Fig. 9DView Fig). Ocellate punctures along the anterior margin of sternites; sixth abdominal sternite lower or depressed than others. Pygidium medial disc finely punctate, delimited by margin that widens towards apical part; basal inner margin rugose.

ELYTRA. Opaque. Interstriae microsculpture shagreened; fine shiny punctures disposed randomly. Striae black, weakly impressed, punctures separated by twice diameter of a puncture on elytral disc.

LEGS. Metacoxa posterior edge bearing row of small impressions, not punctures, separated by twice diameter of the impression. Setigerous punctures next to trochanter insertion larger, twice the size of small ones.

PARAMERES AND LAMELLA COPULATRIX. As in Fig. 9CView Fig, E–F.

Morphological variation

Females ( Fig. 9BView Fig) are distinguished from the males as follows:

HEAD. Surface completely rugose, denser towards anterior portion. Cephalic carina trituberculate, median tubercle higher than lateral tubercles, surface punctate. Lateral tubercles narrow.

PRONOTUM. Weak medial depression, surface punctures greatly impressed separated by length of diameter of a puncture. Medial sulcus conspicuous.

Remarks

This species was first described by Luederwaldt (1931) based on a female as a variety of D. opacipennis  , placing it in the inachus  section as part of the lucasi  group. The holotype locality was Santiago del Estero, Rio Salado, the same later used by Pereira (1941) to described D. bosqi  . The description of the male of this species was by Pereira (1941). Based on the abovementioned taxonomic characters and the possibility of comparing specimens to the holotype, we suggest these species to be synonyms. Within the species group, D. fornicatus  and D. rafanunezi  sp. nov. are the only species where the cephalic carinae of females are trituberculate.

MNHN

France, Paris, Museum National d'Histoire Naturelle

MZSP

Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo

MLP

Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata

CEMT

CEMT

MACN

Argentina, Buenos Aires, Museo Argentina de Ciencias Naturales

BMNH

United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]

MNHN

Museum National d'Histoire Naturelle

MZSP

Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo

MLP

Museo de La Plata

MACN

Museo Argentino de Ciencias Naturales Bernardino Rivadavia

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Dichotomius

Loc

Dichotomius (Luederwaldtinia) fornicatus ( Luederwaldt, 1931 )

Montoya-Molina, Santiago & Vaz-De, Fernando Z. 2021
2021
Loc

Dichotomius bosqi Martínez 1959: 89

Martinez A. 1959: 89
1959
Loc

Dichotomius opacipennis var. fornicatus

Blackwelder R. E. 1944: 207
1944
Loc

Dichotomius bosqi

Pereira F. 1941: 263
1941
Loc

Pinotus opacipennis var. fornicatus

Luederwaldt H. 1931: 302
1931