Dichotomius (Luederwaldtinia) enioi,

Montoya-Molina, Santiago & Vaz-De, Fernando Z., 2021, Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae), European Journal of Taxonomy 734, pp. 1-64: 25-27

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Dichotomius (Luederwaldtinia) enioi

sp. nov.

Dichotomius (Luederwaldtinia) enioi  sp. nov.


Fig. 8View Fig, Map 16


Dichotomius enioi  sp. nov. is separated from other species in the group by the following combination of characters: males interocular space microsculpture shagreened, with flat, well-defined punctures resembling rugosity. Median tubercle conical, twice as high as lateral, with fine apical emargination; lateral tubercles with acute apex. Row of dense uninterrupted ocellate punctures arrange along anterior and posterior margin of Pronotum.Anterior disc declivity with weak anteromedian lobe subemargination. Interstriae microsculpture shagreened.


Dichotomius enioi  sp. nov. is an eponym after Enio Cano, a Guatemalan scientist who contributes to the study of Scarabaeoidea  .

Material examined

Holotype GUATEMALA • ♂; “ Petén, Flores, El Remate , Biot. Cerro Cahuí; 28-29 VII 1995; Col. E.B. Cano ” /“Bosque guamil alto, Colecta nocturna, con heces humanas” / “ Dichotomius agenor (Harold)  , Det. E. Cano, 1996.” [white, p]; CEMT. 


COSTA RICA – Guanacaste • 2 ♂♂, 2 ♀♀; Pq. Nac. Santa Rosa. Parnaud.; CEMT  Puntarenas • 1 ♂, 1 ♀; Est. Queb. Bonita , 50 m. Res. Biol. Carara L-N-194500; 469850; Jun. 1993; J.C. Saborío leg.; CEMT. 

GUATEMALA – Petén • 1 ♂, 1 ♀; Flores, El Remate, Biot. Cerro Cahuí; 28–29 Jul. 1995; E.B. Cano leg.; CEMT  1 ♀; Parque Nac. Tikal , área #1-B; 5–6 Sep. 1995; G. Orellana leg.; CEMT. 

MEXICO – Campeche • 3 ♂♂, 2 ♀♀; Xpujil; 26 Jul. 1995; J.L. Moreno and T. Mestre leg.; CEMT  .



MEASUREMENTS. Length: 21.47 mm. Dorsal and ventral surface black.

HEAD. Anterior surface rugose. Interocular surface microsculpture shagreened, with flat, well-defined punctures. Clypeus with central emargination separating two short, rounded clypeal teeth. Cephalic carina wider than high. Median tubercle twice as high as lateral tubercles, conical in shape and with fine apical emargination, lateral tubercles apical acute. Ventral clypeal process truncate (see Fig. 2BView Fig).

PRONOTUM. Wider than long. Pronotal median disc shiny covered with simple well-impressed punctures, scattered dense uninterrupted ocellate punctures along anterior and posterior margins, absent in lateral margins ( Fig. 8A, CView Fig). Anterior disc pronounced declivity with impressed longitudinal sulcus producing weak anteromedian lobe subemargination. Hypomeral disc smooth, external margin bearing multiple rows of disorganized long setae, approximately same length, (see Fig. 2EView Fig).

ABDOMEN. Prosternum surface microsculpture shagreened. Mesosternum and metepisternum surface covered by large ocellate setigerous punctures. Metasternum setigerous punctures restricted to anterior and lateral edges of mestasternal lobe. Setigerous punctures connecting antero-median edge with lateral through margin of mesocoxa (see Fig. 2GView Fig); lateral edge of metasternum with ocellate setigerous punctures separated by diameter of a puncture, setae reaching metepisternum suture. Sternite with ocellate punctures along anterior margins of each sternite, denser in lateral areas. Pygidium microsculpture shagreened with scattered well-impressed simple punctures; basal margin bearing row of ocellate punctures.

ELYTRA. Interstriae microsculpture shagreened ( Fig. 8CView Fig). Striae with ocellate punctures separated by diameter of a puncture on elytral disc.

LEGS. Fore calcar spiniform, profemur surface covered by clearly impressed punctures ( Fig. 8DView Fig); metacoxa posterior margin bearing row of organized ocellate setigerous punctures.


Morphological variation

Females ( Fig. 8BView Fig)

As males except for the following characters:

HEAD. Surface completely transversely rugose, denser on anterior portion. Cephalic carina quadridentate and greatly elevate, median tubercles higher than lateral.

PRONOTUM. Without medial declivity and sub-emargination, punctures dense and deeply impressed, separated between them by once the diameter of same.

ABDOMEN. Sternites lateral punctuation denser than in males. Pygidium surface punctation wider and denser than in males.


It is possible that Kohlmann & Solis (1997) and López-Guerrero (2005) reported this species for Central America as Dichotomius agenor  .