Dichotomius (Luederwaldtinia) deyrollei ( Harold, 1869 ),

Montoya-Molina, Santiago & Vaz-De, Fernando Z., 2021, Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae), European Journal of Taxonomy 734, pp. 1-64: 22-25

publication ID

https://doi.org/10.5852/ejt.2021.734.1233

publication LSID

lsid:zoobank.org:pub:663A0C2D-E3CF-4B0C-8BA1-566B174829A2

DOI

http://doi.org/10.5281/zenodo.4545849

persistent identifier

http://treatment.plazi.org/id/03850669-FF85-FFD6-FE64-25543A2AC9B3

treatment provided by

Plazi

scientific name

Dichotomius (Luederwaldtinia) deyrollei ( Harold, 1869 )
status

 

Dichotomius (Luederwaldtinia) deyrollei ( Harold, 1869) 

Fig. 7View Fig, Map 5

Pinotus deyrollei Harold, 1869: 139  (original description).

Pinotus deyrollei  – Luederwaldt 1929: 99. — Blackwelder 1944: 69.

Dichotomius deyrollei  – Roze 1955: 44. — Medina et al. 2001: 138. — Sarmiento & Amat 2014: 93–95.

Diagnosis

Dichotomius deyrollei  is separated from the other species in the group by the following combination of characters: clypeus with central emargination displaying two rounded, dorsally angled clypeal teeth. Cephalic carina, higher than wide, antero-posteriorly flattened. Median tubercle approximately twice as high as lateral tubercles and gently inclined anteriorly, apically expanded and emarginate. Anterior portion of pronotum displaying sharp declivity, antero-medial area deeper and convex than lateral and where the longer, median tubercle can fit if head capsules is rotated dorsally.

Material examined

Lectotype (here designated) COUNTRY UNKNOWN • ♂; “Brasilia. P. Deyrollei Harold  ” [white with red margins, hw]/ “Ex- Musaeo E. Harold” [white with black margins, p]/ “Museum Paris ex Coll. R. Oberthur” [green, p]/ “LECTOTYPE ♂ Pinotus deyrollei Harold  des. F.Z. Vaz-de-Mello, 2014” [red with black margin, phw]; MNHN. 

Paralectotype

COUNTRY UNKNOWN • 1 ♀; “Ex-Musaeo E. Harold” [white with black margins, p]/ “Museum Paris ex Coll. R. Oberthur” [green, p]/ “LECTOTYPE ♀ Pinotus deyrollei Harold  des. F.Z. Vaz-de-Mello, 2014” [red with black margin, phw]; MNHN  .

Additional material

COLOMBIA – Casanare • 1 spec.; Yopal, Guarataro; 600 m a.s.l.; Bosque seco Tropical ; Mar. 1996; A. Lopera leg.; ECC  . – Meta • 1 spec.; PNN Tinigua, Varzea ; Mar. 1993; Ivan Jiménez leg.; ECC  . – Norte de Santander • 1 ♂; 3 Km N of Chinacota; 1000 m a.s.l.; dung carrion traps, T10; 10 May 1974; S. Peck leg.; CEMT  1 spec.; Villa Marina ; 1200 m a.s.l.; Feb. 2007; S. Ardila leg.; ECC  . – Santander • 10 specs; Charalá Vda. El Salitre, Fca. Buenos Aires ; 1730 m a.s.l.; 6º32′39″ N, 73º39′6.9″ W; 11 Apr. 2016; C.Giraldo and A. León, J leg.; IAvHGoogleMaps  .

TRINIDAD • 1 ♂; B.W.I Arima Valley ; 30 Apr. 1950; CEMT  1 ♀; 2 May 1953; CEMT  .

VENEZUELA – Mérida • 2 ♀♀; Arzobispo Chacón, Mucutuy ; 1380 m a.s.l.; 8º14′18″ N, 71º17′12″ W; 13 Mar. 2015; Racconcia; CEMTGoogleMaps  . – Miranda • 1 ♂; Guarenas, Curupao ; 19 Jul. 1959; Bordon leg.; MLP  1 ♂; Táchira, 4 Km SE of San Cristobal; 700 m a.s.l.; dung traps; 19 May 1974; S. Peck leg.; CEMT  1 ♂; P. Natural Paramillo ; Jul. 1993; J. Blanco leg.; CEMT  1 ♂; Rio Negro ; 25 Mar. 1984; B. Joffre leg.; CEMT  1 ♂; 2.5 km from San Cristobal to Chorro del Indio; 1100 m a.s.l.; 7 p. m.; 1 Jul. 1989; D. Havranek leg.; BMNH  . – Yaracuy • 2 ♂♂, 1 ♀; Bolivar Aroa ; 710 m a.s.l.; 10º0′0″ N, 68º0′0″ W; heces humanas; 20 Jul. 2009; M. Asmüssen, P. Colmenares and H. Martínez leg.; CEMTGoogleMaps  .

Redescription

Male

MEASUREMENTS. Length: 28.7 mm.

HEAD. Surface of anterior third rugose not reaching base of cephalic carina. Clypeus central emargination separating two rounded and dorsally angled clypeal teeth ( Fig. 7A, DView Fig). Cephalic carina higher than wide, antero-posteriorly flattened; anterior surface with well-marked punctures. Median tubercle approximately twice as high as lateral and gently inclined anteriorly, expanded and emarginate apically. Lateral tubercles, in frontal view, with two acute apices ( Fig. 7DView Fig). Ventral clypeal process truncate (see Fig. 2BView Fig).

PRONOTUM. Wider than long. Pronotal disc shiny and smooth, covered by fine punctures. Antero-medial edge with strong convex declivity, which in the centre could accommodate the large frontal carina ( Fig. 7AView Fig). Row of interrupted ocellate punctures arranged along anterior and posterior margins, fewer and scattered on lateral margins. Hypomeron anterior edge, presenting rounded ocellate punctures with long setae, punctures separated by one time the width of a puncture. Hypomeron medial disc smooth (see Fig. 2EView Fig), without punctures. External margin bearing single row of organized long setae, all approximately of same length (see Fig. 2FView Fig).

ABDOMEN. Mesosternum and metepisternum fully covered by dense ocellate-setigerous punctures. Metasternum setigerous punctures restricted mainly to lateral and anterior areas ( Fig. 7HView Fig). Pilosity of anterior lobe of metasternum connecting with lateral pilosity through margin of mesocoxa ( Fig. 7HView Fig). Lateral surface of metasternum covered by ocellate setigerous punctures separated by twice width of a puncture, setae long and dense, covering lateral surface of metasternum and metepisternal suture. Ocellate setigerous punctures along antero-lateral margins, denser at lateral areas. Pygidium surface covered by fine punctures, basal margin with scattered ocellate punctures.

ELYTRA. Interstriae microsculpture shagreened (see Fig. 2JView Fig). Striae deeply impressed, crenulate, punctures separated by diameter of a puncture on elytral disc.

LEGS. Metacoxa posterior edge bearing single row of disorganized setigerous punctures, separated by half or less the diameter of a puncture.

PARAMERES AND LAMELLA COPULATRIX. As Fig. 7CView Fig, E–F.

Morphological variation

Females ( Fig. 7BView Fig)

As males except for the following characters:

HEAD. Surface transversely rugose, denser on anterior portion. Cephalic carina with pair of high medial tubercles.

PRONOTUM. With weak medial knob, observable in lateral view. Uninterrupted ocellate punctures throughout lateral margins. Hypomeron punctures ocellate and setigerous, equally spaced; setae longer and denser than in males. Metasternum almost completely covered by ocellate setigerous punctures, metasternal lobe smooth with fine and deeply impressed punctures.

ELYTRA. Smooth, with scattered fine punctures.

ABDOMEN. Sternites dense ocellate punctures covering anterior margin.

Remarks

This species occurs in Colombia and Venezuela. The distribution of the species closely follows that of the Andes Mountains. The original description notes that the type specimens were from “ Brésil ”, without any other specification of the locality. However, as far as we know this Andean species does not occur in Brazil.

MNHN

France, Paris, Museum National d'Histoire Naturelle

CEMT

CEMT

MLP

Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata

BMNH

United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]

MNHN

Museum National d'Histoire Naturelle

MLP

Museo de La Plata

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Dichotomius

Loc

Dichotomius (Luederwaldtinia) deyrollei ( Harold, 1869 )

Montoya-Molina, Santiago & Vaz-De, Fernando Z. 2021
2021
Loc

Dichotomius deyrollei

Medina C. A. & Lopera-Toro A. & Vitolo A. & Gill B. 2001: 138
Roze J. A. 1955: 44
1955
Loc

Pinotus deyrollei

Blackwelder R. E. 1944: 69
Luederwaldt H. 1929: 99
1929
Loc

Pinotus deyrollei

Harold E. 1869: 139
1869