Amplirhagada davidsoniana, Köhler, 2011

Köhler, Frank, 2011, Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae), Records of the Australian Museum 63 (2), pp. 167-202 : 177-179

publication ID	https://doi.org/ 10.3853/j.0067-1975.63.2011.1581
publication LSID	lsid:zoobank.org:pub:1BCD4085-D2B9-400D-B504-8C85C30303D6
persistent identifier	https://treatment.plazi.org/id/D59B24C2-EAD8-423E-951F-2148DD2F3523
taxon LSID	lsid:zoobank.org:act:D59B24C2-EAD8-423E-951F-2148DD2F3523
treatment provided by	Felipe
scientific name	Amplirhagada davidsoniana
status	sp. nov.

Treatment

Amplirhagada davidsoniana View in CoL n.sp.

Type locality. Western Australia, NW Kimberley, Admiralty Gulf, 2.6 km E Davidson Point , near Voltaire Passage; 14°14'55"S 125°36'50"E ( RFS 09-2 ; coll. V. Kessner, 9 Jun 1987) ( Fig. 1 View Figure 1 ) GoogleMaps .

Type material. Holotype WAM S34716 View Materials (preserved specimen) (Pl. 1.5; Table 1) . Paratypes AM C.472923 (14 preserved specimens), WAM S34717 View Materials (30 preserved specimens) , FMNH 220282 View Materials (44 preserved specimens), paratypes AM C.472924 (15 shells), WAM S34718 View Materials (36 dried shells) , FMNH 220283 View Materials (54 shells) .

Etymology. In reference to Davidson Point.

Description

Shell (Pl. 1.5; Fig. 14 A–C View Figure 14 ). Broadly conical with moderately elevated spire. Thin to solid, translucent. Periphery evenly rounded to slightly angulate; upper and basal sectors of whorls well rounded. Umbilicus 50–80 percent concealed by columellar reflection. Background colour horn to yellowish brown; peripheral band absent to well developed, usually diffuse, rather thin, brown, visible on last whorls only; subsutural band absent or diffuse and very thin, brown, visible on last whorls only; ventral and outer lip colour horn; inner lip translucent, pale white. Protoconch c. 2.4 mm in diameter, comprising about one whorl, with well-developed, radially elongated pustulations. Teleoconch with fine, regular axial growth lines, rounded in cross-section; spaces equal to thickness of lines; sculpture evenly distributed across whorls of shell and across whorl diameter, height of lirae reduced underneath suture. Angle of aperture less than 45 degrees; outer lip sharp to moderately thick, rounded, expanded, slightly reflected; basal node absent or very weak. Parietal wall of inner lip inconspicuous. Average shell size 13.1±0.8 × 18.3± 0.9 mm ( Table 1).

Radular and jaw morphology ( Fig. 14 D–G View Figure 14 ). Tooth formula C + 12–14 + 4 + 24–28, with 150–160 rows of teeth (n = 2). Jaw with 11 plates.

Genital morphology ( Figs 15–16 View Figure 15 View Figure 16 ). Penis straight, tubular, slightly longer than anterior part of oviduct. Penial sheath very delicate. Length of penial retractor muscle equivalent to length of penial complex. Penial verge elongately conical, with pointed tip, extending about 1/5 of length of penial chamber. Inner penial wall entirely covered by well-developed pustules arranged in longitudinal rows or pilasters. Main stimulatory pilaster relatively short, coneshaped comprising proximal third of inner penial wall; with lateral corrugations. Vas deferens very thick, not undulating, entering penial sheath within proximal third of penial complex. Vagina long, tubular; inner vaginal wall with rows of laterally expanded pustules that are densely arranged in longitudinal rows. Rows of pustules giving rise to corrugated pilasters at proximal end of vagina. Inner wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix slightly extending base of spermoviduct. Free oviduct short, comprising less than half of length of anterior part of oviduct. Spermoviduct about longer than anterior part of oviduct.

Aestivation strategy: Rock sealer.

Remarks. Anatomical description based on dissections of three specimens. Listed by Solem (1991) as “ Amplirhagada NSP 21”. Shell similar to A. imitata ( Smith, 1894) , A. solemiana Köhler, 2010 and A. indistincta Köhler, 2010 , but differing by shell size, distinct corrugation of inner penial wall and much longer vagina.