Amplirhagada coffea, Köhler, 2011

Köhler, Frank, 2011, Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae), Records of the Australian Museum 63 (2), pp. 167-202 : 186-187

publication ID	https://doi.org/ 10.3853/j.0067-1975.63.2011.1581
publication LSID	lsid:zoobank.org:pub:1BCD4085-D2B9-400D-B504-8C85C30303D6
persistent identifier	https://treatment.plazi.org/id/0F68235C-52D0-4A69-9E9C-114C3F50CE20
taxon LSID	lsid:zoobank.org:act:0F68235C-52D0-4A69-9E9C-114C3F50CE20
treatment provided by	Felipe
scientific name	Amplirhagada coffea
status	sp. nov.

Treatment

Amplirhagada coffea View in CoL n.sp.

Type locality. Western Australia, NW Kimberley, George Water, 16 km SW of Mt. Grey , near Barlee Impediment; 15°49'10"S 124°36'00"E (RFS-23-3; coll. V. Kessner, 23 Jun 1987) ( Fig. 1 View Figure 1 ) GoogleMaps .

Type material. Holotype WAM S34731 View Materials (preserved specimen) (Pl. 1.10; Table 1) . Paratypes AM C.472932 (3 preserved specimens, WAM S34732 View Materials (8 preserved specimens), FMNH 220945 View Materials (13 preserved specimens), AM C.472933 (6 dried shells), WAM S34733 View Materials (10 dried shells), FMNH 220948 View Materials (16 dried shells).

Etymology. Species epithet refers to the dark coffee-brown colour of the shell.

Description

Shell (Pl. 1.10; Fig. 29 A–C View Figure 29 ). Semi-globose, with rather high spire; solid. Periphery compressed to angulate; upper sector slightly shouldered, basal sector rounded. Umbilicus completely concealed by columellar reflection. Background colour crème to brown; sub-sutural and mid-whorl bands usually blend into each other covering entire shell surface in chestnut brown; ventral and outer lip colour light brownish horn; inner lip whitish. Shell surface glossy. Protoconch 3.3 mm in diameter, comprising about 1.2 whorls, with indistinct radially elongated pustulations. Teleoconch with well-developed, regular axial growth lines, evenly distributed across shell surface. Angle of aperture 30–45 degrees; outer lip rounded to slightly angulate, slightly expanded, well reflected, basal node of lip present, palatal node absent. Average shell size 17.9±1.6 × 21.4± 1.1 mm ( Table 1).

Genital morphology ( Fig. 30 View Figure 30 ). Penis straight, more or less of same length as anterior part of oviduct. Penial sheath proximally thick, distally delicate. Length of penial retractor muscle equivalent to about a third of length of penial complex. Penial verge slender, moderately long, extending about 1/6 of length of penial chamber. Distal part of inner penial wall entirely covered by regular pustulation; pustules relatively large, conical. Proximal part of inner penial wall covered by many irregular, corrugated pilasters formed from fused rows of pustules. No main stimulatory pilaster present. Vas deferens entering penial sheath within distal quarter of penial complex. Rest of genital system unknown.

Aestivation strategy. Rock sealer.

Remarks. Description based on dissection of one specimen. Listed by Solem (1991) as “ Amplirhagada NSP 27” together with dry shells found on Boongaree Island, which were tentatively considered conspecific as based on similar shell. The species from Boongaree was previously described as A. boongareensis Köhler, 2010 . Amplirhagada coffea differs from the former by usually having a concealed umbilicus, different shell and inner lip colour, glossy surface, strongly reflected outer lip. Species with similar “bee-hive” shaped shells, such as A. boongareensis , A. decora Köhler, 2010 and A. vialae differ by having large cone-shaped main pilasters and distinct pustulation of inner penial wall. Material from a second sampling site at the mainland coast (RFS-26–3, Brecknock Harbour, opposite Camden Island) is tentatively being considered conspecific for its similar shell and penial anatomy.