Amplirhagada bendraytoni, Köhler, 2011

Köhler, Frank, 2011, Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae), Records of the Australian Museum 63 (2), pp. 167-202 : 191

publication ID	https://doi.org/ 10.3853/j.0067-1975.63.2011.1581
publication LSID	lsid:zoobank.org:pub:1BCD4085-D2B9-400D-B504-8C85C30303D6
persistent identifier	https://treatment.plazi.org/id/0384503A-FFA3-3C46-FC00-FE79FADC698E
treatment provided by	Felipe
scientific name	Amplirhagada bendraytoni
status	sp. nov.

Treatment

Amplirhagada bendraytoni View in CoL n.sp.

Type locality. Western Australia, NW Kimberley,York Sound, 4 km SE of Cape Brewster , S of Coronation Island, between beach and base of escarpment; 15°07'40"S 124°56'15"E ( RFS 12-3 ; coll. V. Kessner, 12 Jun 1987) ( Fig. 1 View Figure 1 ) GoogleMaps .

Type material. Holotype WAM S34740 View Materials (preserved specimen) (Pl. 1.13; Table 1) . Paratypes WAM S34741 View Materials (2 preserved specimens) , FMNH 220448 View Materials (3 preserved specimens) , WAM S34742 View Materials (3 dried shells) , FMNH 2200447 View Materials (5 dried shells) .

Etymology. Named in honour of my friend and neighbour Ben Drayton from Sydney.

Description

Shell (Pl. 1.13; Fig. 37 A–B View Figure 37 ). Broadly conical to semiglobose with moderately elevated spire. Solid, not translucent. Periphery angulate; upper and basal sectors rounded. Umbilicus 50–80% concealed by columellar reflection. Background colour horn with a yellowish brown tone; spiral bands absent; outer and inner lip colour lighter than shell. Protoconch not differentiable on SEM micrograph. Teleoconch with very faint axial growth lines; evenly distributed across shell surface. Angle of aperture c. 45 degrees; outer lip moderately thick, well rounded, slightly expanded, not reflected; basal node absent. Parietal wall of inner lip absent. Average shell size 15.2±1.5 × 21.3± 1.3 mm ( Table 1).

Radular and jaw morphology ( Fig. 37 C–G View Figure 37 ). Radular Tooth formula C + 13–18 + 4 + 18–22, with 130–140 rows of teeth (n = 2). Jaw with nine plates.

Genital morphology ( Figs 38–39 View Figure 38 View Figure 39 ). Penis straight, tubular, thin, as long as anterior part of oviduct. Penial sheath thick. Penial retractor half as long as penis. Penial verge tiny, with pointed tip. Inner penial wall entirely smooth, with several rather indistinct longitudinal pilasters. Main stimulatory pilaster short, at proximal half of inner penial wall, corrugated by lateral lamellae, each lamella supporting usually three little hooks that are arranged into longitudinal rows that run along the central portion of the pilaster. Vas deferens thin, entering penial sheath close to penial apex, forming an extended loop. Vagina very short, proximally inflated; inner vaginal wall and wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix slightly extending base of spermoviduct. Free oviduct comprising more than half of length of anterior part of oviduct. Spermoviduct longer than anterior part of oviduct.

Aestivation strategy. Free sealer.

Remarks. Anatomical description based on dissections of two specimens. Listed by Solem (1991) as “ Amplirhagada NSP 31”. The combination of an almost entirely smooth inner penial wall with a sort main pilaster, a tiny verge and a thick penial sheath are peculiar to only this species.