Cephennodes (Cephennodes) pinnapes, Jałoszyński, Paweł, 2015

Jałoszyński, Paweł, 2015, The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 4033 (3), pp. 393-410 : 400-401

Cephennodes (Cephennodes) pinnapes sp. n.

( Figs 6–8 , 17–18 )

Type material. Holotype: CHINA (Sichuan Province): ♂, four labels: " CHINA: W-Sichuan / Ya'an Pref., Tianquan Co. / Jiajin Shan, valley above Labahe" [white, printed]; "N.R. Station, 57kmW Ya'an / 30.06.63 N, 102.25.18E, light forest / 1800m, 12.VII.1999, leg.A.Pütz" [white, printed]; "Sammlung / Andreas Pütz / Eisenhüttenstadt" [yellow, printed]; " CEPHENNODES (s. str.) / pinnapes m. / det. P. Jałoszyński, '15 / HOLOTYPUS " [red, printed] (cAP). Paratype: 1 ♂, same data as holotype (cPJ).

Diagnosis. Abdomen of male strongly modified, with sternite V bearing median lamina strongly projecting ventrally; elytral subhumeral lines much shorter than half EL; aedeagus in ventral view with the distal margin of apical projection perpendicular to the long axis of aedeagus and slightly recurved.

Description. Body of male ( Fig. 6 ) slightly flattened, oval with feebly marked constriction between pronotum and elytra, moderately dark brown with slightly lighter legs and distinctly lighter palps, covered with light brown vestiture. BL 1.09–1.19 mm.

Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL 0.14 mm, HW 0.30 mm; vertex and frons confluent and weakly convex, supraantennal tubercles distinct. Punctures on frons and vertex very fine but dense, separated by spaces subequal to diameters of punctures; setae sparse, short, suberect. Antennae moderately long, four terminal antennomeres distinctly and gradually broadening, AnL 0.60–0.61 mm, antennomeres I–II elongate, III slightly transverse, IV–VI each about as long as broad, VII slightly elongate, VIII– X each slightly transverse, XI about as long as IX–X together, twice as long as broad, with rounded, blunt apex.

Pronotum semioval, convex at middle and slightly flattened near hind corners, broadest between middle and posterior third but only slightly narrowing posteriorly; PL 0.33–0.35 mm, PW 0.48–0.50 mm. Anterior margin slightly arcuate; lateral margins strongly rounded in anterior half and nearly straight in posterior third; hind pronotal corners nearly right-angled; posterior margin shallowly bisinuate; lateral carinae indistinct, not delimited from disc and not thickened; lateral antebasal pits shallow but distinct, each nearly equally distant from posterior and lateral pronotal margin. Punctures on median part of pronotal disc as fine and dense as those on frons and vertex, punctures becoming denser, but not larger or deeper, toward lateral pronotal margins. Setae on pronotal disc dense and moderately long, suberect.

Elytra slightly more convex than pronotum, oval, broadest between middle and anterior third; EL 0.63–0.70 mm, EW 0.53–0.59 mm, EI 1.19; each subhumeral line marked as sharp border between more convex humeral and less convex adsutural region, as long as only 0.32 EL, lines weakly divergent caudad; basal elytral fovea on each elytron located closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, rounded together. Punctures on elytra as fine and dense as those on pronotum; setae dense, moderately long, suberect. Hind wings well-developed, about twice as long as elytra.

Metaventrite with large and deep postmesocoxal impressions.

Abdomen ( Figs 7–8 ) modified, sternite V with median lamina strongly protruding ventrally with lateral pair of large cavities at base of lamina.

Legs moderately long and slender; pro- and mesotibiae distinctly recurved, metatibiae with large apical broadening.

Aedeagus ( Figs 17–18 ) simonis form, AeL 0.15 mm; median lobe in ventral view drop-shaped with apical portion subtriangular; apical projection short and broad, with distal margin perpendicular to the long axis of aedeagus and slightly recurved; parameres slender, each with one apical and one subapical seta.

Female. Unknown.

Distribution. Central-southern China: Sichuan Province.

Etymology. The specific epithet pinnapes means "winged on the feet, having winged feet" and refers to the distal broadening and flattening of the metatibiae.

Remarks. Within the inflatipes species group, the Vietnamese C. patellatus and C. pseudopatellatus have strikingly long elytral subhumeral lines (about as long as 0.7 EL), while in all Chinese species, including the newly described C. pinnapes , the subhumeral lines are as long as only about 0.3 EL. Males of C. inflatipes and C. kopeipes differ from C. pinnapes in an unmodified abdomen. The three species with modified abdomen, C. abdominalis , C. parabdominalis and C. pinnapes can be distinguished on the basis of body proportions and, first of all, the aedeagi (examined in strictly ventral view). The distal margin of the apical projection in C. abdominalis is evenly convex; in C. parabdominalis it is nearly straight; in C. pinnapes the margin is weakly recurved. The apex of apical projection in C. abdominalis is triangular and pointed, whereas apices in C. parabdominalis and C. pinnapes are rounded and blunt. The distal margin of the apical projection in C. abdominalis and C. parabdominalis is oblique in relation to the long axis of the aedeagus, forming with this axis about 45º-angle, whereas in C. pinnapes the apical margin is perpendicular to the long axis of the aedeagus. Moreover, the apex of median lobe in ventral view in C. abdominalis and C. parabdominalis forms a projection parallel to the long axis of the aedeagus, whereas in C. pinnapes the apex of median lobe is protruding obliquely outward from the longitudinal axis.

Figures

FIGURES 1 – 10. Dorsal habitus of holotype males (1, 4, 6, 9, 10), elytral apices in dorsal (2) and posterodorsal (3) views; head of male in dorsolateral view (5); abdomen of male in ventral (7) and lateral (8) views. Cephennodes (s. str.) intricatus sp. n. (1 – 3); Cephennodes (s. str.) monofoveatus sp. n. (4 – 5); Cephennodes (s. str.) pinnapes sp. n. (6 – 8); Cephennodes (s. str.) elegantissimus sp. n. (9); Cephennodes (s. str.) palpalis sp. n. (10). Abbreviations: ai, apical impression; csf, circumsutural flattening; csr, C-shaped ridge; sb, setal brush; s 3 – 8, abdominal sternite III – VIII.

FIGURES 11 – 18. Aedeagus in ventral (11, 13, 15, 17) and lateral (12, 14, 16, 18) views. Cephennodes (s. str.) intricatus sp. n. (11 – 12); Cephennodes (s. str.) cryptointricatus sp. n. (13 – 14); Cephennodes (s. str.) monofoveatus sp. n. (15 – 16); Cephennodes (s. str.) pinnapes sp. n. (17 – 18).