Orizabus delgadoi Ratcliffe and Cave, 2010

Ratcliffe, Brett C. & Cave, Ronald D., 2010, New species of Orizabus Fairmaire (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) from Mexico and Guatemala, with a revised key and checklist of the species in the genus, Insecta Mundi 2010 (127), pp. 1-16 : 4-7

Orizabus delgadoi Ratcliffe and Cave , new species

(Fig. 4-6, 26, 28)

Type Material. Holotype male labeled “ MEX:

Oaxaca / Cerro Zempoaltepetl/ 3000 m, 20-22 vii

09/ Curoe, Ramirez col”. Two paratypes with same

data. Holotype deposited at the University of Ne-

braska State Museum (Lincoln, NE), one paratype

in the Leonardo Delgado collection (Xalapa, Mexico),

and one paratype in the B.C. Ratcliffe collection

(Lincoln, NE).

Holotype. Male. Length 22.1 mm; width 13.0 mm.

Color dark reddish brown, almost black. Head:

Surface completely, coarsely, transversely rugose.

Frontoclypeal region with transverse, arcuate ca-

rina with low tubercle at middle. Clypeus with

subapical carina bilobed, reflexed. Interocular

width equals 7.5 transverse eye diameters. An-

tenna 10-segmented, club distinctly longer than

segments 2-7. Pronotum: Surface on anterior half

with dense, moderately large punctures, becom-

ing rugopunctate on sides and angles; posterior

half sparsely punctate, punctures small. Fovea and

tubercle absent. Posterior border lacking marginal

bead. Elytra: Surface with 5 furrowed, punctate

striae between suture and median edge of hu-

merus, punctures moderately large, ocellate. In-

terval 2 with short row of similar punctures at

base. Sides with similar, but less distinct, rows of

punctures. Pygidium: Surface punctate; punctures

moderate in density and size, shallow. Apex with

marginal bead and short, transverse, preapical

depression at center. In lateral view, surface regu-

larly convex. Legs: Protibia lacking teeth on ex-

ternal margin. Metatibiae at apex with 23 and 26

spinules. Venter: Prosternal process long, colum-

nar, apex rounded, densely covered with long, red-

dish brown setae. Parameres: Each paramere with

small, subapical notch on lateral edge, apices, in

lateral view, not concave (Fig. 5-6).

Variation. Male paratypes (2). Length 19.9-20.5

mm; width 12.2-12.5 mm. The paratypes do not

differ significantly from the holotype. The puncta-

tion on the pygidium is reduced in one specimen

(punctures are more shallow) and nearly obsolete

in the other specimen.

Etymology. We take great pleasure in dedicating

this species to Leonardo Delgado (Instituto de Figure 4-6. Orizabus delgadoi . 4) Dorsal view. 5-6) Ecología, Xalapa, Mexico) in recognition of his fine Parameres, caudal and lateral views.

work documenting the dynastine fauna of Mexico.

11

Holotype. Male. Length 19.6 mm; width 12.1 mm. Color dark reddish brown. Head: Surface vaguely, coarsely, transversely rugose to rugopunctate (surface appears worn). Frontoclypeal region with transverse carina, carina becoming tuberculiform at middle. Clypeus with subapical carina weakly bilobed, reflexed. Interocular width equals 5.0 transverse eye diameters. Antenna 10-segmented, club slightly longer than segments 2-7. Pronotum: Surface with moderately dense, moderately large punctures except along midline on basal half of center of disc where punctures sparse, small; anterior angles coarsely rugopunctate; broad band along lateral margins with dense, moderately large punctures. Fovea absent. Etymology. The specific epithet for this species is derived from the Spanish word mezcla, meaning a mixture or medley. This species is so named in reference to its mixture of character states seen in species of Orizabus .

Distribution. Orizabus mezclus is known from a single specimen from the Guatemalan department of Suchitepéquez, near Patulul ( Fig. 26 ).

Temporal Distribution. June (1).

Diagnosis. Orizabus mezclus superficially resembles O. clunalis and O. cuernavacensis Delgado and Deloya in the form of the edentate protibia, sculpturing of the pronotum, and parameres. Unlike O. clunalis , however, it does not have a fovea on the pronotum, and each paramere has a small tooth on the ventrolateral edge ( Fig. 12 ), whereas there is only an angular swelling in O. clunalis ( Fig. 19 ). It differs from O. cuernavacensis because it is larger (nearly 20 mm versus 17 mm), and it has a distinctive, transverse groove just before the marginal bead of the pygidium ( Fig. 11 ), whereas there is no groove in O. cuernavacensis .

Biology. Nothing is known of the life history of this species. It was collected at lights at an elevation of 1,399 m.

Figures

Figure 26. Distribution map for the new Orizabus species described in this paper.

Figure 12-13. Orizabus mezclus, Parameres, caudal and lateral views..

Figure 17-25. Parameres of Orizabus spp. 17) O. batesi. 18) O. brevicollis. 19) O. clunalis. 20) O. cuernavacensis. 21) O. fairmairei. 22) O. pyriformis. 23) O. rubricollis. 24-25) O. teamscaraborum.

Figure 10-11. Orizabus mezclus. 10) Dorsal habitus. 11) Oblique view of left pygidial apex.