Agathidium invisitatum Miller and Wheeler, 2005

MILLER, KELLY B. & WHEELER, QUENTIN D., 2005, Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part Ii. Coleoptera: Leiodidae, Bulletin of the American Museum of Natural History 2005 (291), pp. 1-167 : 110-112

Agathidium invisitatum Miller and Wheeler , new species Figures 138 , 240–243 , 365

TYPE MATERIAL: Holotype, Ƌ in FMNH labeled ‘‘ MEXICO: Mexico W slope Nevado de Toluca, 11,000 ft ix.16.1973/under pine bark A. Newton/ HOLOTYPE Agathidium invisitatum Miller and Wheeler, 2003 [red label with black line border]’’.

TYPE LOCALITY: Mexico, State of Mexico, west slope of Nevado de Toluca, 11,000̍.

DIAGNOSIS: This species is identifiable by the presence of a prominent series of four denticles on the male metafemur with the largest at about two­thirds the distance from the base of the metafemur (fig. 138). The metasternum is moderately broad medially and the eyes are moderately large and finely faceted. The male genitalia are distinctive with a long and broad operculum that is apically distinctly emarginate (fig. 240). The rami of the operculum are apically expanded ventrad (fig. 242).

DESCRIPTION: Body moderately large (TBL = 3.28–3.82 mm), broad (PNW/TBL = 0.44–0.52), robust, rounded, strongly contractile.

Head and pronotum dark yellow­red; elytra dark yellow­red, distinctly iridescent; venter yellow­red; antennae and palpi yellow­red; legs yellow.

Head broad (MDL/OHW = 0.53–0.63), dorsal surface flattened, dorsoventrally compressed; with very fine punctures, each with a short, very fine seta, surface between punctures shiny, very lightly microreticulate; frontoclypeal suture obsolete medially; eyes prominent, not strongly compressed; gula flat; antennae moderately long (ratios: length I:II:III = 1.9:1.0:2.0, width VII:VIII:IX = 1.1:1.0:2.0). Pronotum very large, broad (PNL/PNW = 0.72–0.78), strongly convex, anterolateral lobes strongly produced, lateral margin broadly curved, not angulate; with fine, sparse punctures, each with a short, very fine seta, surface between punctures shiny, smooth. Elytra broad, lateral margins strongly rounded, apically rounded (SEL/ELW = 0.93–1.07); punctation and surface similar to pronotum; sutural stria absent. Flight wings strongly reduced. Mesosternum moderately broad, not declivitous; medial carina low but distinctly developed. Metasternum narrow (MTL/MTW = 0.13–0.17), slightly concave medially, distinctly dorsally sloped anteriorly; oblique femoral carinae moderately well developed, forming a low carina medially.

Male tarsi 5­5­4; pro­ and mesobasotarsomeres somewhat laterally expanded, protarsomeres more so and with small field of ventral spatulate setae; mandibles not modified; metafemur moderately broad, with several prominent, small teeth medially along posterior margin (fig. 138); metasternal fovea small with small pencil of fine setae. Median lobe in lateral aspect slender basally, more robust submedially, with small, but prominent, lobelike lateral carinae between which fit the lateral lobes, apical portion relatively short, curved ventrad, apex pointed (fig. 242); in ventral aspect moderately slender, apical portion narrowed, slender to expanded, rounded apex (figs. 240, 241); operculum in lateral aspect sender, apically expanded and slightly curved ventrad (fig. 242), in ventral aspect broad, short, apically narrowly emarginate, each ramus with small lateral tooth (fig. 240); lateral lobes slender, curved basally, apical one­third broader and sinuate, apices rounded, each with 2 stout setae (fig. 243).

Female not examined.

ETYMOLOGY: Named for the Latin word invisitatus, meaning ‘‘strange’’, for the unusual teeth on the male metafemur.

DISTRIBUTION: This species has been collected from central Mexico (fig. 365).

PARATYPE: MEXICO: San Luis Potosi: 14 mi W Xilitla, 29 Jun 1973, 4800̍, under bark, Liqidambar forest, A Newton (1, FMNH).

DISCUSSION: The species is known from under bark at 4800 to 11,000 ft elevation.

Figures

Figs. 118–144. Agathidium concinnum­ and A. oniscoides­group species, left male metafemur, ventral aspect: 118, A. concinnum. 119, A. kimberlae. 120, A. vaderi. 121, A. bituberculum. 122, A. oedema. 123, A. gomezae. 124, A. hidalgoense. 125, A. skoliosternum. 126, A. erythromelas. 127, A. rhamphastes. 128, A. megoniscoides. 129, A. grumum. 130, A. triangularum. 131, A. lobosternum. 132, A. potosii. 133, A. popocatepetlae. 134, A. hyle. 135, A. stenomma. 136, A. tribulosum. 137, A. tribulograndum. 138, A. invisitatum. 139, A. multidentatum. 140, A. sejunctum. 141, A. grandidentatum. 142, A. andersoni. 143, A. disgregum. 144, A. oaxacaense. Bars = 0.5 mm.

Figs. 232–260. Agathidium oniscoides­group species, aedeagus: 232–235, A. tribulosum: 232, ventral; 233, median lobe, apex, ventral; 234, median lobe, lateral; 235, left lateral lobe, lateral. 236–239, A. tribulograndum: 236, ventral; 237, median lobe, apex, ventral; 238, median lobe, lateral; 239, left lateral lobe, lateral. 240–243, A. invisitatum: 240, ventral; 241, median lobe, apex, ventral; 242, median lobe, lateral; 243, left lateral lobe, lateral. 244–247, A. multidentatum: 244, ventral; 245, median lobe,

Figs. 365, 366. Geographic distribution of Agathidium oniscoides­group species: 365, A. andersoni = -; A. disgregum = ^; A. invisitatum = •; A. oaxacaense = *. 366, A. impensum = *; A. oculeum = •; A. recurvatum = -; A. cheneyi = ^.