Lepanus pygmaeus (Macleay, 1888)

Gunter, Nicole L. & Weir, Thomas A., 2019, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group, Zootaxa 4564 (1), pp. 41-80 : 49-50

publication ID

https://doi.org/10.11646/zootaxa.4564.1.2

publication LSID

lsid:zoobank.org:pub:27F70B5F-24DD-4C9E-BBA7-2B87476BD49C

DOI

https://doi.org/10.5281/zenodo.5922792

persistent identifier

https://treatment.plazi.org/id/03832713-FFA3-8D7B-A3DA-FBC050F297A8

treatment provided by

Plazi

scientific name

Lepanus pygmaeus
status

 

Key to species Lepanus pygmaeus species group

Due to the subtle nature of some diagnostic characters, some specimens are difficult to diagnose. Images of some diagnostic characters are provided in Figs. 4–12 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 11 View FIGURE 12 to help facilitate diagnoses. Given the small size of specimens (1.8–3.0 mm), a high-powered microscope with adequate lighting is recommended for identification. Furthermore, caution must be taken when interpreting setation on specimens as fine setae rub off and may appear absent.

1 Protibia with three teeth on outer edge, basal tooth sometimes very small ( Fig. 4 View FIGURE 4 I–J)................................ 2

- Protibia with two teeth on outer edge ( Fig. 4 View FIGURE 4 G–H, K–L)....................................................... 3

2 Pygidium with a tear-drop shaped depression extending under the usually straight upper edge, depression with an elongate tubercle ( Fig. 9E View FIGURE 9 ); male metatibiae with almost straight inner edge; female 6 th abdominal ventrite swollen in middle; aedeagus ( Fig.12D View FIGURE 12 )......................................................... Lepanus sauroni Gunter & Weir , new species

- Pygidium with an almost ovoid depression just extending under the usually angulate upper edge, depression with a round tubercle ( Fig. 9A View FIGURE 9 ); male metatibiae with expanded inner edge, creating a distinct flange ( Fig. 4 View FIGURE 4 D–E); female 6 th abdominal ventrite not swollen; aedeagus as in Fig. 11E View FIGURE 11 .............................. Lepanus pecki Gunter & Weir , new species

3 Front edge of protibia truncate ( Figs. 3G View FIGURE 3 , 4 View FIGURE 4 G–H); apical digit absent in females ( Fig. 4G View FIGURE 4 )............................. 4

- Front edge of protibia deeply angulate ( Fig. 3H View FIGURE 3 ); apical digit present in females (as in Fig. 4I View FIGURE 4 )......................... 5

4 Pygidial depression widely pear shaped with a somewhat triangular tubercle in the upper third, upper edge of pygidium narrowly angulate, depression just extending under the edge ( Fig. 9D View FIGURE 9 ); punctures along basal margin of prothorax slightly elongate, differing from those on the disc; dorsal part of the eye moderate in size; aedeagus ( Fig. 12C View FIGURE 12 ).............................................................................................. Lepanus pygmaeus ( Macleay, 1888)

- Pygidial depression round with a round tubercle in the upper third, upper edge of pygidium broadly angulate, depression extending under the edge ( Fig. 9F View FIGURE 9 ); punctures along basal margin of prothorax more round, similar to those on the disc; dorsal part of the eye narrower in size; aedeagus ( Fig. 12E View FIGURE 12 )..................... Lepanus tozerensis Gunter & Weir , new species

5 Pygidial depression narrow and vertically elongate, without a tubercle, occupying less than half the area of the pygidial disc ( Fig. 8F View FIGURE 8 ), basal carina of head complete, with a slight medial angle ( Fig. 4B View FIGURE 4 ); aedeagus as in Fig. 11A View FIGURE 11 ..................................................................................... Lepanus lentil Gunter & Weir , new species

- Pygidial depression of different shape, usually occupying at least one third to half the area of the pygidial disc (smaller and round in L. menendezae ), depression with or without a distinct tubercle; basal carina of head either obsolete, or interrupted medially............................................................................................. 6 6 Pygidial depression a narrow or broad U-shape, without a distinct tubercle ( Figs. 8A, G View FIGURE 8 ); basal carina of head absent or obsolete; basal margin of pronotum slightly angulate medially...................................................... 7

- Pygidial depression different, either round (e.g., Figs. 8C, I View FIGURE 8 ), ovoid (e.g., Fig. 8H View FIGURE 8 ), pear or tear-drop shaped (e.g., Figs. 8D View FIGURE 8 , 9 View FIGURE 9 B–C), with a distinct tubercle; basal carina of head present, medially interrupted ( Fig. 4A View FIGURE 4 ); basal margin of pronotum rounded medially............................................................................................. 8

7 Pygidial depression narrow U-shape, reticulate, occupying the medial third of the pygidial disc ( Fig. 8A View FIGURE 8 ); surface of the head finely reticulate between clypeogenal suture and anterior edge, rest of head smooth (as in Fig. 4A View FIGURE 4 ); dorsal part of eye moderate, interocular space about 10 times the width of dorsal part of the eye; with a fine carina extending backwards from the mesometaventral suture ( Fig. 4C View FIGURE 4 ); elytral surface reticulate in apical half and at the sides, striae slightly impressed, striae 5–7 similar to striae 1–4; aedeagus as in Fig. 10A View FIGURE 10 .......................... Lepanus andersonorum Gunter & Weir , new species

- Pygidial depression broad U-shape, smooth, occupying at least medial half of the pygidial disc ( Fig. 8G View FIGURE 8 ); surface of head entirely smooth; dorsal part of eye narrow in size, interocular space about 16 times the width of dorsal part of the eye; without a fine carina extending backwards from the meso-metaventral suture; elytral surface smooth, striae superficial, striae 5–7 weaker than striae 1–4; aedeagus as in Fig. 11B View FIGURE 11 .......................... Lepanus lingziae Gunter & Weir , new species

8 Pygidial depression pear or tear-drop shaped ( Figs. 8D View FIGURE 8 , 9 View FIGURE 9 B–C).................................................. 9

- Pygidial depression round or ovoid ( Figs. 8 View FIGURE 8 B–C, E, H–I)..................................................... 11

9 Pygidium with an evident secondary basal line across base of depression, pygidial depression tear-drop shaped, not reaching base, tubercle slightly elongate ( Fig. 9C View FIGURE 9 ); mesoventrite with discernible punctures at least at sides; surface of the head finely reticulate between clypeogenal suture and anterior edge (as in Fig. 4A View FIGURE 4 ); aedeagus as in Fig. 12B View FIGURE 12 ...................................................................................... Lepanus pungalina Gunter & Weir , new species

- Pygidium with secondary basal line not evident, merged with base, pygidial depression reaching base, tubercle elongate or an inverted V ( Figs. 8D View FIGURE 8 , 9B View FIGURE 9 ); mesoventrite with or without discernible punctures; surface of head entirely smooth.......... 10

10 Pygidial depression widely pear shaped, tubercle elongate ( Fig. 9B View FIGURE 9 ); elytral flange and groove relatively wide; colour dark brown to black with red humeri ( Fig. 6E View FIGURE 6 ); mesoventrite with discernible punctures at least at sides; aedeagus as in Fig. 12A View FIGURE 12 ................................................................. Lepanus podocarp Gunter & Weir , new species

- Pygidial depression narrowly tear-drop shaped, tubercle an inverted V ( Fig. 8D View FIGURE 8 ); elytral flange and groove narrower; colour uniformly dark brown ( Fig. 5D View FIGURE 5 ); mesoventrite virtually impunctate; aedeagus as in Fig. 10D View FIGURE 10 ............................................................................................ Lepanus gubara Gunter & Weir , new species

11 Pygidial depression small and round, occupying less than half of the pygidial disc, separated from base, tubercle round and centrally located; secondary basal line evident ( Fig. 8I View FIGURE 8 ); aedeagus as in Fig. 11D View FIGURE 11 ................................................................................................. Lepanus menendezae Gunter & Weir , new species

- Pygidial depression larger, occupying about half of the pygidial disc, usually reaching the base ( Figs. 8 View FIGURE 8 B–C, E, H); secondary basal line evident or not............................................................................... 12

12 Pygidium with evident secondary basal line across base of depression, depression usually not reaching base, tubercle tubular with a dark tip ( Fig. 8E View FIGURE 8 ); mesoventrite with discernible punctures at least at sides and often reticulate; surface of the head finely reticulate between clypeogenal suture and anterior edge; elytral striae 6 and 7 appearing double, similar to striae 1–5; male metatibia with expanded inner edge, creating a small flange ( Fig. 4F View FIGURE 4 ); aedeagus as in Fig. 10E View FIGURE 10 .................................................................................... Lepanus guthrieae Gunter & Weir , new species

- Pygidium with secondary basal line not evident, merged with base, upper edge of depression angulate, tubercle of various shapes ( Figs. 8 View FIGURE 8 B–C, H); mesoventrite virtually impunctate, not reticulate; surface of head usually entirely smooth; elytral striae 6 and 7 appearing single, different from striae 1–4 or 1–5; male metatibia with inner edge not expanded............... 13

13 Apical tooth on protibiae with a slight notch on the inner edge ( Fig. 4K View FIGURE 4 ) [often best viewed from below]; tubercle within pygidial depression low, round, or slightly elongate, same colour as depression ( Fig. 8B View FIGURE 8 ); eye canthus not dividing the eye (as in Fig. 3F View FIGURE 3 ); aedeagus as in Fig. 10B View FIGURE 10 .................................. Lepanus crenidens Gunter & Weir , new species

- Apical tooth on protibiae without a notch on the inner edge; tubercle within pygidial depression with a darker tip, either shaped as an inverted V or slightly horizontal; eye canthus nearly dividing the eye (as in Fig. 3E View FIGURE 3 )........................... 14

14 Pygidial depression with tubercle somewhat inverted V shaped ( Fig. 8C View FIGURE 8 ); elytral striae 5–7 with discernible punctures at least in basal half; elytral flange and groove relatively wide; basal margin of pronotum with a distinct row of punctures ( Fig. 5C View FIGURE 5 ); aedeagus as in Fig. 10C View FIGURE 10 .............................................. Lepanus feehani Gunter & Weir , new species

- Pygidial depression with tubercle slightly horizontal, not like an inverted V ( Fig. 8H View FIGURE 8 ); elytral striae 5–7 without discernible punctures; elytral flange and groove narrower; basal margin of pronotum without a distinct row of punctures ( Fig. 6B View FIGURE 6 ); aedeagus as in Fig. 11C View FIGURE 11 ................................................ Lepanus mckenziei Gunter & Weir , new species

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Lepanus