Styringomyia rose MARTIN, THEISCHINGER & BILLINGHAM, 2018

Theischinger, Günther, D, Zacariah & Martin, John, 2018, The genus Styringomyia LOEW in Australia (Diptera: Tipuloidea: Limoniidae), Linzer biologische Beiträge 50 (2), pp. 1587-1633 : 1600-1601

Styringomyia rose MARTIN, THEISCHINGER & BILLINGHAM nov.sp. ( Figs 45-47 )

T y p e m a t e r i a l. Holotype ♂: Australia, Northern Territory, Koongarra, 15 km E of Mt Cahill, 15-xi-1972, D.H. Colless (ANIC). Paratypes: 3♀♀, same data as holotype (ANIC).

D e s c r i p t i o n

Male ( Figs 45, 46 )

Head. Including rostrum, palps and antennae dark brownish grey to black, a fine pale grey midline in posterior half.

Thorax. Brownish grey to black. Base of coxae and trochanters of fore- and midleg greyish brown, approximately basal half of femora and very base of tibiae paler; coxa and trochanter of hindleg dark yellow, remainder missing.

Wings. Hyaline without infumation or clouds. Halters grey.

Abdomen. Tergites brown to black, sternites pale to dark brown.

Hypopygium. Tergite 9/10 largely parallel sided, apically tongue-shaped. Sternite 9 apically narrow, ending in two long digitate lobes, each with apical, subapical and basal seta, the lobes separated by U- to V-shaped notch. Basal two thirds of gonocoxites stout, apical third digitate with apical spine moderately thick and slightly more than half as long as gonocoxites. Gonostyli complex, including a wide, apically trifid element with two branches pointed and the third whip-like with subapical seta, and at least two, possibly three, comb-like elements of different size and shape, partly toothed, partly setose. What is detectable in ventral view of the aedeagus distinctly trilobed.

Dimensions. Wing length 3.3 mm.

Female ( Fig. 47 )

Head, Thorax and Abdomen. Coloration generally somewhat paler than in male; midline of head less distinct; hind leg similar to fore- and mid-leg; abdomen pale to medium brown.

Terminalia. Cerci apparently bifurcate with each lobe bearing an apical spine and reaching backward approximately as far as the tips of the apically slightly sinuously curved hypogynial valves.

Dimensions. Wing length 2.6-3.0 mm.

D i s t r i b u t i o n. Northern; known only from Northern Territory ( Map 4 ).

E t y m o l o g y. This species is named for Rose, wife of John Martin, junior author of the present paper.

D i s c u s s i o n. The male of Styringomyia rose nov.sp. shares a bifid apex of sternite 9, and complex gonostyli with the species of the S. bancrofti group. In S. rose , however, the base of the gonocoxites is stout and the apical section is digitate with a moderately thick spine versus more conical with mesal expansion and thicker spine. Also, the aedeagus is much more prominent (trilobed) in S. rose than in the species of the S. bancrofti group. Styringomyia rose may well be the sister taxon of the S. bancrofti group.

Figures

Map 4: Distribution of the ungrouped Styringomyia species: S. rose, S. flagro, S. naso, S. danbulla, S. kimberley.

Figs 45-47: Styringomyia rose nov.sp., terminalia: (45, 46) male: (45) ventral; (46) dorsal; (47) female, lateral.