Ochodaeus, Dejean, 1821

Paulsen, M. J., 2019, New species of Ochodaeus Dejean from Madagascar II (Coleoptera: Scarabaeoidea: Ochodaeidae), Insecta Mundi 706 (706), pp. 1-10 : 8-9

Updated key couplets to Ochodaeus spp. in Madagascar

The following couplet changes to the species key in Paulsen (2019) allow for determination of all Madagascan species having a granulate pronotum, including the three newly described species.

2(1). Elytral intervals with setose tubercles of 2 sizes, with setae of larger tubercles flattened, scale-like........................................................ O. pygmaeus Paulian — Elytral intervals with tubercles either absent or of uniform size and without flattened setae. 3

3(2). Clypeus with median tubercle or small horn that is distinct from anterior margin ( Fig. 4–6 )........................................................................... 4 — Clypeus lacking free tubercle or horn, anterior margin usually thickened medially ( Fig. 7–8 ). 6

4(3). Mentum with a knobby protuberance on each side ( Fig. 1d ).... O. meridialis Paulsen , n. sp. — Mentum lacking protuberances laterally............................................ 5

5(4). Clypeus ( Fig. 5 ) with median tubercle anterior to frontoclypeal suture; anterior margin of clypeus indistinct and eroded medially........................ O. umbonulus Paulsen , n. sp.

— Clypeus ( Fig. 6 ) with small horn located on frontoclypeal suture; anterior margin of clypeus distinct and entire throughout.......................... O. iniquipes Paulsen , n. sp.

6(3). Median thickening of clypeal bead variable, lateral thickening always evident ( Fig. 7 ); body length usually> 9.0 mm........................................ O. isoanalensis Paulian

— Median thickening of clypeal bead simply convex, bead slender laterally ( Fig. 8 ); body length usually 4.5–8.0 mm............................................ O. miliaris Klug

Figures

Figure 1. Ochodaeus meridialis Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view.c) Right stridulatory peg, dorsal view (elytra lifted). d) Head, female paratype, ventral view. Arrows indicating knob-like protuberances of mentum. e) Holotype labels.

Figures 4–8. Key characters of Madagascar Ochodaeus species. 4) Head of O. meridialis Paulsen, n. sp., showing central tubercle of clypeus. 5) Head of O. umbonulus Paulsen, n. sp., showing tubercle anterior to straight frontoclypeal suture. 6) Head of O. iniquipes Paulsen, n. sp., showing minute horn on posteriorly curved frontoclypeal suture. 7) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 8) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin.