Cloeodes espinillo, Nieto, Carolina & Richard, Barton, 2008

Nieto, Carolina & Richard, Barton, 2008, The genus Cloeodes (Ephemeroptera: Baetidae) in Argentina with new generic synonymy and new species, Zootaxa 1727, pp. 1-21 : 8-11

publication ID

https://doi.org/ 10.5281/zenodo.181238

DOI

https://doi.org/10.5281/zenodo.6233769

persistent identifier

https://treatment.plazi.org/id/036D87E7-D532-FFCD-FF6C-B7E4FDDBFEA3

treatment provided by

Plazi

scientific name

Cloeodes espinillo
status

sp. nov.

Cloeodes espinillo View in CoL n. sp.

( Figs. 15–30 View FIGURES 15 – 19 View FIGURES 20 – 30 )

Male imago ( Fig. 15 View FIGURES 15 – 19 ). Length: body, 6.4–6.5 mm. Fore wings, 5.9–6.0; hind wings 1.1–1.2 mm. Head yellowish brown, compound eyes pale yellow, height of stalk half length of eye diameter. Antennae pale yellow. Thorax yellowish brown, sterna yellowish brown. Legs pale yellow. Leg I: tibia 1.5 times the length of femur, tarsi as long as tibia with 4 segments decreasing in length apically. Fore wings ( Fig. 16 View FIGURES 15 – 19 ) hyaline, except costal and subcostal spaces translucent. Hind wings ( Figs. 17 View FIGURES 15 – 19 a and b) present, 0.2 times the length of fore wing; costal projection pointed and located in the basal third of anterior margin. Posterior margin of the metanotum strongly projected dorsoventrally ( Fig. 18 View FIGURES 15 – 19 ). Abdomen pale yellow. Genitalia ( Fig. 19 View FIGURES 15 – 19 ) with bases of forceps close together, segment III rounded. Cerci pale yellow.

Female imago. Length: body, 6.6–6.7 mm. Fore wings, 7.2 mm; hind wings, 1.2 mm. Similar to male imago except thorax yellow. Abdomen yellow with tracheae pigmented.

Nymph ( Fig. 20 View FIGURES 20 – 30 ). Length: body, 5.5–6.0 mm; cerci, 2.8–3.0 mm; terminal filament, 2.5 mm. Head yellowish brown, longer than wide; compound eyes of the male yellowish brown. Antennae pale yellow, 3 times the head capsule; pedicel 1.5 times the length of scape. Mouthpart: labrum ( Fig. 21 View FIGURES 20 – 30 ) with one subapical seta centrally and two setae near lateral margin, anterior margin with basally bifid setae near midline and apically bifid setae near lateral margin. Left mandible ( Fig. 22 View FIGURES 20 – 30 ) without setae between prostheca and mola, thumb of molar area transverse to anterior margin. Right mandible without setae between prostheca and mola (similar to Fig. 58 View FIGURES 55 – 67 ). Hypopharynx ( Fig. 23 View FIGURES 20 – 30 ): lingua with a rounded projection. Maxillae (similar to Fig. 60 View FIGURES 55 – 67 ), palpi as long as galea-lacinia, two segmented, segment II with a constriction. Labium ( Fig. 24 View FIGURES 20 – 30 ) with segment III of palpi rounded.

Thorax yellowish brown ( Fig. 20 View FIGURES 20 – 30 ). Fore wing pads pale yellow. Sterna pale yellow. Legs ( Fig. 25 View FIGURES 20 – 30 ) yellow. Dorsal edge of femora with a row of pointed spines, apically with a subquadrangular projection with a pair of blunt spines ( Fig. 26 View FIGURES 20 – 30 ). Tarsi with a row of short spines. Tarsal claws 0.4 times the length of tarsi. Hind wing pads present.

Abdomen yellowish brown, segments II–III and VI with brown spots as in Fig. 20 View FIGURES 20 – 30 , segment IX–X brown, posterior margin of terga with spines ( Fig. 27 View FIGURES 20 – 30 ). Sterna pale yellow, segments II–VI with a small tuft of long fine setae laterally on each side ( Fig. 28 View FIGURES 20 – 30 ). Paraprocts with spines apically (similar to Fig. 51 View FIGURES 43 – 51 ). Gills translucent white, 2 times the length of each tergum, main trachea pigmented ( Fig. 29 View FIGURES 20 – 30 ). Caudal filaments yellow, apical third yellowish brown, with flattened setae basally sclerotized ( Fig. 30 View FIGURES 20 – 30 ). Cerci ( Fig. 30 View FIGURES 20 – 30 ) with long spines toward external margin every two segments, terminal filament with long spines toward midline dorsally and Etymology. Espinillo is the name of one of the rivers where this species was collected.

Diagnosis. Cloeodes espinillo n. sp. can be distinguished from the other species of the genus by the following combination of characters. In the male imago, 1) hind wings 0.2 times the length of the fore wings ( Figs. 16, 17 View FIGURES 15 – 19 a); 2) hind wings ( Fig. 17 View FIGURES 15 – 19 b) with costal projection located in the first third of the anterior margin; 3) genitalia ( Fig. 19 View FIGURES 15 – 19 ) with bases of forceps close together, segment III rounded. In the nymph, 1) hind wing pads present; 2) lingua with a rounded projection ( Fig. 23 View FIGURES 20 – 30 ); 3) dorsal edge of femora with a row of pointed spines, apically with a subquadrangular projection with a pair of blunt spines ( Fig. 26 View FIGURES 20 – 30 ); 4) abdominal color pattern yellowish brown with segments II–III and VI with brown spots as in Fig. 20 View FIGURES 20 – 30 , segments IX–X brown; 5) caudal filaments with flattened setae basally sclerotized ( Fig. 30 View FIGURES 20 – 30 ), cerci with long spines toward external margin every two segments and terminal filament with long spines toward midline dorsally and ventrally every two segments (similar to Fig. 14 View FIGURES 5 – 14 ).

Material. Holotype: male nymph: ARGENTINA, Córdoba, Río Los Espinillos, 13–16/ IX/ 1997, Nieto & Molineri colls. Paratypes: 4 nymphs, 5 male imagos (reared) and 16 male imagos, same data as holotype. 18 nymphs: Depto. Pocho, arroyito afluente Río Pocho, S 31°13'17'', W 65°08'26'', 1030 m, 14/ XI/ 2001, Orce & Nieto colls. Material housed in IFML.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Ephemeroptera

Family

Baetidae

Genus

Cloeodes

GBIF Dataset (for parent article) Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF